Imshaugia aleurites (Ach.) S.L.F. Mey.

Mycologia, 77: 337, 1985. Basionym: Lichen aleurites Ach. - Lichenogr. Suec. Prodr.: 117, 1799.
Synonyms: Cetraria aleurites (Ach.) Th. Fr.; Imbricaria aleurites (Ach.) DC.; Parmelia aleurites (Ach.) Ach.; Parmeliopsis aleurites (Ach.) Nyl.; Parmeliopsis pallescens (Hoffm.) Hillmann; Parmeliopsis placorodia sensu Jatta non auct.; Platysma diffusum (Weber) Nyl.
Distribution: N - Frl, Ven (Nimis 1994, Nascimbene & Caniglia 2000b, 2002c, 2003c, Nascimbene & al. 2006e, Nascimbene 2008c), TAA (Nascimbene & Caniglia 2000b, 2002c, Caniglia & al. 2002, Nascimbene & al. 2005, 2006, 2006e, 2007b, 2008b, 2008c, 2009, 2010, 2014, 2022, Nascimbene 2006b, 2006c, 2013, 2014, 2014c, Lang 2009, Watson 2014, Nascimbene & Marini 2015, Nimis & al. 2015, Trindade & al. 2021), Lomb (Alessio & al. 1995, Nascimbene & al. 2006e, Gheza 2019), Piem (Isocrono & al. 2004, Isocrono & Piervittori 2008), VA (Piervittori & Isocrono 1999, Matteucci & al. 2008, Isocrono & al. 2008), Emil (Tretiach & al. 2008, Fariselli & al. 2020), Lig (Giordani & al. 2009). C - Tosc (Benesperi 2007, Benesperi & al. 2007, Tretiach & al. 2008, Brackel 2015). S - Cal (Puntillo 1996, Incerti & Nimis 2006), Si.
Description: Thallus foliose, heteromerous, dorsiventral, adnate, forming orbicular to irregular (2-)3-5(-7) cm wide rosettes. Lobes 1-4 mm long, 0.5-1.5(-3) mm wide, flat, linear to sublinear, divergent to contiguous, irregularly branched, with rounded, often forked tips. Upper surface whitish grey to bluish grey, sometimes wrinkled or shallowly pitted in older parts, shiny at the tips of lobes, with abundant, laminal, cylindrical, simple to branched, to c. 2 mm tall, fragile isidia that with age tend to break and coalesce into a granulose mass. Lower surface white to tan, somewhat wrinkled, with brownish, simple, c. 0.5-1 mm long rhizines. Upper cortex paraplectenchymatous, of tightly packed, anticlinally oriented hyphae, with a pored epicortex, the cell walls with Cetraria-type lichenan; medulla white. Apothecia rare, lecanorine, 2-7 mm across, with a convex to flat, brown, epruinose disc and a persistent, often isidiate thalline margin. Epithecium brown; hymenium and hypothecium colourless. sci 8-spored, clavate, very thin-walled, with a K/I+ blue, tall tholus penetrated by a faintly amyloid apical cushion, the wall K/I-, surrounded by a blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid, (5-)6-7(-9) x (3.5-)4-5(-6) µm. Pycnidia rare, black, mostly marginal, clearly projecting. Conidia ellipsoid to dumbbell-shaped, 3-4(-5) x c. 1 µm, Psora-type, borne terminally from joints of conidiogenous hyphae. Photobiont chlorococcoid. Spot tests: upper cortex and medulla K+ deep yellow, C-, KC-, P+ deep yellow-orange, or reddish, UV-. Chemistry: upper cortex with atranorin and chloroatranorin; medulla with thamnolic acid (major) and decarboxythamnolic acid (trace).
Note: a circumboreal-montane to cool-temperate species found on acid bark, mostly of conifers and on decorticated stumps, with optimum near treeline; common only in the Alps, much rarer in the mountains of Southern Italy.
Growth form: Foliose, narrow lobed
Substrata: bark and lignum
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: very common
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[20369], U.S.A. Carolina Borealis, Grandville County, Creedmore, ad truncum Pini sp.. Leg. W.L. Culberson, 03.1955, det. W.L. Culberson. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1967



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34870)
2002/02/17
center of thallus with isidia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (9123)
2001/11/25
detail of isidia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34870)
2002/02/17
peripheral lobes



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (9123)
2001/11/25


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php



Felix Schumm - CC BY-SA 4.0
[19051], Australia, Victoria, Mount Delegate, 8 km south-south-west of Delegate (NSW), 37°07' S, 148°54' E, 1300 m , growing on dead wood in Eucalyptus pauciflora dominated woodland. Leg. J.A. Elix (20019) & H. Streimann, 10.04.1985.



Felix Schumm - CC BY-SA 4.0
[19051], Australia, Victoria, Mount Delegate, 8 km south-south-west of Delegate (NSW), 37°07' S, 148°54' E, 1300 m , growing on dead wood in Eucalyptus pauciflora dominated woodland. Leg. J.A. Elix (20019) & H. Streimann, 10.04.1985.



Felix Schumm - CC BY-SA 4.0
[19051], Australia, Victoria, Mount Delegate, 8 km south-south-west of Delegate (NSW), 37°07' S, 148°54' E, 1300 m , growing on dead wood in Eucalyptus pauciflora dominated woodland. Leg. J.A. Elix (20019) & H. Streimann, 10.04.1985.


Felix Schumm - CC BY-SA 4.0
: [19051], Australia, Victoria, Mount Delegate, 8 km south-south-west of Delegate (NSW), 37°07' S, 148°54' E, 1300 m , growing on dead wood in Eucalyptus pauciflora dominated woodland. Leg. J.A. Elix (20019) & H. Streimann, 10.04.1985
th: thamnolic acid



Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
August 2017
as v. tilesii



Felix Schumm - CC BY-SA 4.0
[20369], U.S.A. Carolina Borealis, Grandville County, Creedmore, ad truncum Pini sp.. Leg. W.L. Culberson, 03.1955, det. W.L. Culberson. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1967



Felix Schumm - CC BY-SA 4.0
[19051], Australia, Victoria, Mount Delegate, 8 km south-south-west of Delegate (NSW), 37°07' S, 148°54' E, 1300 m , growing on dead wood in Eucalyptus pauciflora dominated woodland. Leg. J.A. Elix (20019) & H. Streimann, 10.04.1985.



Felix Schumm - CC BY-SA 4.0
[20369], U.S.A. Carolina Borealis, Grandville County, Creedmore, ad truncum Pini sp.. Leg. W.L. Culberson, 03.1955, det. W.L. Culberson. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1967


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38801



P.L. Nimis CC BY-SA 4.0
TSB 37321



P.L. Nimis CC BY-SA 4.0
TSB 37321



P.L. Nimis CC BY-SA 4.0
TSB 37322



P.L. Nimis CC BY-SA 4.0
TSB 37322



P.L. Nimis CC BY-SA 4.0
TSB 38801



Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0
TSB 37321



Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0
TSB 37321



Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0
TSB 37321



Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0
TSB 37322



Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0
TSB 37322


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=359&lang=en
France, Doubs