Umbilicaria brigantium Zschacke

Verh. bot. Ver. Prov. Brandenburg, 69: 14, 1927.
Synonyms: Lasallia brigantium (Zschacke) Llano
Distribution: C - Tosc, Sar (Nöske 2000). S - Cal (Lich. Ital. Exs. 33: Isocrono & al. 2019).
Description: Thallus foliose-umbilicate, usually monophyllous, 2-8 cm in diam., grey to white, attached by a stout, central holdfast. Upper surface with numerous conspicuous, convex, round to oval pustules, areolate. Isidia scarce, marginal or laminal, black, irregularly branched but not coralloid. Lower surface black, markedly areolate, erhizinate. Upper cortex paraplectenchymatous, with a layer or dark-walled cells close to the photobiont layer; epicortex discontinuous, of hyaline, large, thin-walled hyphae; medulla composed of an up to 45 μm thick upper, arachnoid layer with hyphae encrusted with crystals, and a lower layer of thick-walled hyphae; lower cortex of periclinally arranged, thin-walled hyphae, up to 150 μm thick, with a strongly necrotic lower zone. Apothecia common, 1-2.5(-3) mm across, sessile or immersed, with a black, flat, smooth disc and a persistent, smooth, ecorticate proper margin. Proper exciple forming a palisade plectenchyma with the outer half of necrotic cells, without an arachnoid medullary layer. Asci 1(-2)-spored, elongate-clavate, thick-walled, with a K/I+ blue apical dome. Ascospores muriform, at first hyaline, then pale to dark brown, 50-75 x 22-35 μm. Pycnidia black, immersed. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-; medulla K-, C+ red, KC+ red, P-, UV-. Chemistry: medulla with gyrophoric acid.
Note: on basic siliceous rocks, apparently restricted to Sardinia, Corsica, Tuscany and adjacent islands, always at low altitudes and mostly near the coast.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: very rare
Dry mediterranean belt: extremely rare

pH of the substrata:

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(info)

Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples
Index Fungorum GBIF

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse



Pier Luigi Nimis
Italy
TSB 41767



P.L. Nimis CC BY-SA 04
TSB 13764


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (8218)
2001/11/25


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_brigantium.htm
France, 18/10/2012 - Lumio Arinella Puntu Carchincu, alt. 15 m - Corse


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio
lower surface


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio
apothecium


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=949&lang=en
France, Corse, Lumio