Umbilicaria hispanica (Frey) Davydov, Peršoh & Rambold

Taxon, 67, 1: 221, 2018. Basionym: Umbilicaria brigantium var. hispanica Frey - Ber. schweiz. bot. Ges., 59: 443, 1949.
Synonyms: Lasallia brigantium var. hispanica (Frey) Llano; Lasallia hispanica (Frey) Sancho & A. Crespo; Umbilicaria hispanica (Frey) Davydov, Peršoh & Rambold
Distribution: S - Bas (Potenza & al. 2014), Cal (Puntillo 1996, Potenza & al. 2011), Si (CLU 4848).
Description: Thallus foliose-umbilicate, usually monophyllous, 190-300 μm thick, 2-8 cm in diam., grey to white, attached by a stout, central holdfast. Upper surface with numerous, conspicuous, convex, circular to oval pustules, the central parts echinate-areolate, the margins becoming more or less eroded. Isidia scarce, marginal or laminal, black, irregularly branched but not coralloid. Lower surface creamy grey or brown, blackish around the umbilicus, smooth or slightly areolate, erhizinate. Upper cortex 30-40 μm thick, paraplectenchymatous, with a layer or dark-walled cells close to the photobiont layer; epicortex discontinuous, of hyaline, large, thin-walled hyphae; medulla composed of an up to 25 μm thick upper, arachnoid layer with hyphae encrusted with crystals, and a lower layer of thick-walled hyphae; lower cortex of periclinally arranged, thin-walled hyphae, up to 30 μm thick, with a layer of slightly darkened cells. Apothecia common, 1-2.5(-3) mm across, stipitate, with a black, flat, smooth disc and a persistent, smooth, corticate proper margin. Proper exciple 30-50 μm thick, persistent, dark in outer rim, colourless within, paraplectenchymatous, composed of mesodermatous or pachydermatous cells arranged in a more or less regular palisade, with an arachnoid medullary layer. Asci 1(-2)-spored, elongate-clavate, thick-walled, with a K/I+ blue apical dome. Ascospores muriform, at first hyaline, then pale to dark brown, 40-65 x (17-)20-25 μm. Pycnidia black, immersed. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-; medulla K-, C+ red, KC+ red, P-, UV-. Chemistry: medulla with gyrophoric acid.
Note: on wind-exposed basic siliceous rocks wetted by rain, but avoiding seepage tracks, mostly in upland areas of the Mediterranean Region.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: very rare
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019



P.L. Nimis CC BY-SA 4.0
TSB 11886



P.L. Nimis CC BY-SA 04
TSB 11886



P.L. Nimis CC BY-SA 04
TSB 11886