Peltigera membranacea (Ach.) Nyl.

Bull. Soc. Linn. Normandie, sér. 4, 1: 74, 1887. Basionym: Peltidea canina var. membranacea Ach. - Lichenogr. Univ.: 518, 1810.
Synonyms: Peltigera canina var. membranacea (Ach.) Duby
Distribution: N - VG, Frl (Tretiach & Molaro 2007, Hafellner 2021), Ven (Caniglia & al. 1999, Nascimbene & Caniglia 2000), TAA (Nascimbene 2008b), Lomb (Gheza & al. 2022), Piem (Morisi & Sereno 1995, Isocrono & al. 2004, Isocrono & Piervittori 2008), Emil, Lig (Ravera & al. 2019a). C - Tosc (FI 294), Abr, Sar (Vitikainen 1994). S - Pugl, Bas, Cal, Si.
Description: Thallus foliose, broad-lobed, heteromerous and dorsiventral, grey to brownish grey when dry, bluish grey or blackish green when wet, loosely attached, forming 10-20(-35) cm wide rosettes. Lobes elongate, flattened, contiguous, 1-3(-4) cm broad, up to 10 cm long, often slightly bullate, glabrescent and somewhat shiny towards centre, arachnoid-tomentose and matt at margins, which are usually down-turned and entire. Lower surface whitish, with white to pale brown, conspicuously erect-tomentose (also in thallus centre), raised, 0.5-0.7 mm broad veins, polygonal interstices, and dispersed, slender, 5-7(-13) mm long rhizines with short ramifications perpendicular to the main axis (bottle-brush-like). Upper cortex pseudoparenchymatous; medulla white; lower cortex absent. Apothecia frequent, saddle-shaped, on short, ascending lobes, up to 6 mm in diam, with a brown to reddish brown disc. Paraphyses simple, distinctly thickened above. Asci 8-spored, Peltigera-type, fissitunicate, the thickened apex with a K/I+ blue ring. Ascospores 3(-5)-septate, hyaline to very pale brown at maturity, acicular, thin-walled, (40-)50-65(-80) x 2.5-5 µm. Pycnidia dark, immersed. Conidia fusiform. Photobiont: cyanobacterial (Nostoc, the cells not in long chains). Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: on mossy rocks and at the base of boles in old woodlands, usually on base-rich substrata, with optimum in the montane belt.
Growth form: Foliose, broad lobed
Substrata: soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: rather rare
Oromediterranean belt: absent
Montane belt: rare
Submediterranean belt: rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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(info)

Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples
Index Fungorum GBIF

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (21206)
2001/12/05
lower surface



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31502)
2003/03/03



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31502)
2003/03/03
young and old rhizines


Andres Saag; Owner: University of Tartu



E. Pittao
TSB 35802)



E. Pittao
TSB 35802)



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41952


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_membr.htm
France, bord d'un ruisseau - session AFL 2007 - Finistère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_membr.htm
France, bord d'un ruisseau - session AFL 2007 - Finistère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_membr.htm
France, bord d'un ruisseau - session AFL 2007 - Finistère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_membr.htm
France, bord d'un ruisseau - session AFL 2007 - Finistère


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Huelgoat


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Huelgoat


Joël Querellou - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Huelgoat


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Stangala


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Stangala


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Brennilis


Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Forêt du Cranou


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Cap de la Chèvre


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=420&lang=en
France, Hôpital-Camfrout


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=870&lang=en
France, Brennilis
saxicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=870&lang=en
France, Brennilis
saxicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=870&lang=en
France, Brennilis
saxicolous