Arthonia subvarians Nyl.

Flora, 51: 345, 1868.
Synonyms:
Distribution: N - TAA (Brackel 2016), Lomb (Brackel 2013, 2016), Piem (Brackel 2016). C - Tosc (Brackel 2016), Pugl (Brackel2016).
Description: Thallus inapparent, not lichenized, the hyphae developing inside the hymenia of Lecanora-species, especially of the L. polytropa-group. Apothecia arthonioid, black, 0.12-0.15 mm across, with a slightly convex disc covered with a hyaline gelatinous sheet, and without a proper margin. Proper exciple poorly developed, not evident; epithecium grey-brown, K-; hymenium colourless, I+ deep wine-red, K/I+ blue; paraphysoids septate, branched, c. 2 μm thick, apically swollen and pigmented; hypothecium reddish brown, K+ pale olive brown. Asci (4-)8-spored, broadly clavate to subglobose, semi-fissitunicate, with a large, non-amyloid apical dome, and a distinct ocular chamber, Arthonia-type. Ascospores 1-septate, hyaline, ellipsoid to slightly soleiform, 8.5-12 x 4-5 μm, indistinctly halonate. Photobiont absent. Spot tests: K-, C-, KC-, P-. Chemistry: without lichen substances.
Note: a rarely recorded parasite in the hymenia of Lecanora species, especially of the L. polytropa-group. It was also reported from Toscana and Trentino-Alto Adige (as Arthonia clemens on Protoparmeliopsis muralis, Nimis & al. 1990, BSM 2006-2012), but these finds should be reinvestigated, as the hosts (Protoparmeliopsis muralis, L. varia) leave some doubts about the identity.
Growth form: Lichenicolous fungus
Substrata: rocks
Reproductive strategy: mainly sexual
paras Lecanora spp.

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: very rare
Oromediterranean belt: very rare
Montane belt: very rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Pier Luigi Nimis CC BY-SA 4.0
TSB 42613



Pier Luigi Nimis CC BY-SA 4.0
TSB 42611



Pier Luigi Nimis CC BY-SA 4.0
TSB 42611



Pier Luigi Nimis CC BY-SA 4.0
TSB 42611



Pier Luigi Nimis CC BY-SA 4.0
TSB 42613



Pier Luigi Nimis CC BY-SA 4.0
TSB 42613



Pier Luigi Nimis CC BY-SA 4.0
TSB 42611