Rinodina milvina (Wahlenb.) Th. Fr.

N. Acta Reg. Soc. Sci. Upsal., ser. 3, 3: 224, 1861. Basionym: Parmelia milvina Wahlenb. in Ach. - Meth. Lich. Suppl.: 34, 1803.
Synonyms: Lecanora milvina (Wahlenb.) Ach.; Lecanora sophodes var. scopulina Nyl.; Lecanora sophodes var. submilvina Nyl.; Lecanora subconfragosa Nyl.; Rinodina milvina var. karelica Räsänen; Rinodina milvina var. scopulina (Nyl.) H. Olivier; Rinodina sophodes f. saxicola Kernst.; Rinodina sophodes var. scopulina (Nyl.) Croz.; Rinodina subconfragosa (Nyl.) Flagey
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, TAA (Nascimbene 2003, Nascimbene & al. 2022), Lomb, Piem (Isocrono & al. 2004, Favero-Longo & al. 2015), VA (Piervittori & Isocrono 1999, Matteucci & al. 2008c, 2015c, Favero-Longo & Piervittori 2009, Isocrono & al. 2008), Emil (Tretiach & al. 2008, Fariselli & al. 2020), Lig (Brunialti & al. 1999). C - Tosc, Sar (Nöske 2000, Rizzi & al. 2011, Giordani & al. 2013). S - Pugl, Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si.
Description: Thallus crustose, episubstratic, rimose-areolate, often delimited by a black prothallus, the areoles up to 0.5-1 mm wide, smooth to rugose, grey-brown, reddish brown or dark brown, dull. Apothecia lecanorine, adnate to subsessile, contiguous, sometimes angular by mutual compression, 0.3-1(-1.5) mm across, with a dark brown to black, concave to flat, rarely slightly convex disc, and a thick, smooth, persistent thalline margin. Thalline exciple 60-100 µm wide laterally, corticate; proper exciple colourless, 5-20 µm wide laterally, expanding to 10-40 µm in upper part; epithecium reddish brown; hymenium colourless, (65-)100-120 µm high; paraphyses 1.5-2.5 µm thick at mid-level, not conglutinate, the apical cells 3.5-5.5 µm wide, immersed in dispersed pigment; hypothecium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, often constricted at septum, brown, broadly ellipsoid, (13-)16-19(-23) x (7-)9-11(-13) µm, Milvina-type, often grading into Pachysporaria-type at maturity, with a well-developed torus and ornamented walls, the ontogeny of type A (apical wall thickening after septum formation). Pycnidia immersed. Conidia bacilliform, 4-5 µm long. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a cool-temperate to arctic-alpine, circumpolar lichen found on boulders of base-rich to weakly calciferous siliceous rocks, usually on steeply inclined surfaces, often (but not always) parasitic on other crustose lichens; the Italian distribution ranges from the Alps to the high Mediterranean mountains, with optimum near or above treeline.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
paras crustose lichens when young

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: rather common
Montane belt: rather rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Pier Luigi Nimis - CC BY-SA 4.0
TSB 42444



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34912)
2002/02/21



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20168)
2001/11/29


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm



P.L. Nimis CC BY-SA 4.0
TSB 42444



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34912)
2002/02/21


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R2/Rinodina_milvina.htm
France, 2/9/2014 - Bonneval-sur-Arc, Pont de L'Oulietta, alt. 2500 m - Savoie


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R2/Rinodina_milvina.htm
France, 2/9/2014 - Bonneval-sur-Arc, Pont de L'Oulietta, alt. 2500 m - Savoie