Usnea glabrescens var. fulvoreagens Räsänen

Ann. Acad. Sci. Fenn., A, 34, 4: 20, 1931.
Synonyms: Usnea fulvoreagens (Räsänen) Räsänen
Distribution: N - Frl, Ven, Lomb (Alessio & al. 1995), Emil (Fariselli & al. 2020). C - Umb (Panfili 2000, Ravera & al. 2006), Sar (Zedda 2002, 2002b). S - Pugl (Thüs & Licht 2006).
Description: Thallus fruticose-filamentous, greenish, shrubby, branching mainly isotomic-dichotomously with divergent branches, 3-5(-7) cm long. Main branches with many ramifications along their whole length, cylindrical, <1 mm thick, with conspicuous annular cracks exposing the medulla, not constricted at ramification points, the base black, without longitudinal cracks. Lateral branches not constricted at attachment point. Papillae and tubercles rare to numerous, most frequent on the main branches; fibrils usually numerous, arranged in a fishbone-like pattern, most frequent in the basal parts, 2-4(-10) mm long. Soralia rounded and discrete, deeply excavate, often reaching down to the central axis, occupying more than half the diameter of the branch, originating from cortex; isidiomorphs always absent. Cortex thick (10-13% of total thickness); medulla homogeneous, loose to compact, thin (12-26%), white; central axis white, rather thick (30-50%). Apothecia not observed in Italian material. Photobiont chlorococcoid. Spot tests and chemistry: cortex with usnic acid, K-, C- KC-, P; medulla with different chemotypes: 1) with norstictic and stictic acids, and variable amounts of diffractaic acid (K+ pale yellow to reddish orange, P+ yellow to orange), by far the most frequent chemotype; 2) with norstictic and squamatic acids (K+ red, P+ orange); 3) without medullary substances (K-, P-). A further chemotype with salazinic and norstictic acid (K+ red, P+ yellow to orange), plus different accessory substances, has been reported from Scandinavia.
Note: on twigs and branches of conifers, more rarely of deciduous trees in cold-humid, open woodlands with frequent fog; chemically heterogeneous. Some Italian records could refer to U. perplexans. U. glabrescens s.lat. seems to be polyphyletic (see Mark & al. 2016), and needs further integrative studies.
Growth form: Fruticose filamentous
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: University of Tartu
Estonia.
Herbarium: TU (31444)
2007/04/26
Bracelet-like soralia of U. fulvoreagens


Tiiu Tõrra; Owner: University of Tartu
Estonia.
Herbarium: TU (31444)
2007/04/26
Branches of U. fulvoreagens


Andres Saag; Owner: University of Tartu


Andres Saag; Owner: University of Tartu


Triin Aimla; Owner: University of Tartu


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia


Tiiu Tõrra; Owner: Tiiu Tõrra - Institute of Ecology and Earth Sciences, University of Tartu, Estonia