Description: Primary thallus squamulose, persistent and forming cushions, the squamules 4-8 mm long, 1-2 mm wide, upcurled and deeply incised, brownish green above, yellowish white beneath, the base somewhat darker and very finely arachnoid. Podetia not always present, stunted, hollow inside, dark brownish green, up to 3(-5) cm tall, terminated by a up to 4 mm wide cup which usually proliferates from the center (1-2 tiers), the surface areolate-corticate, esorediate, the base strongly melanotic. Apothecia rare, brown. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, pyriform, with a colourless jelly, developing at the margins of cups. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P+ bright yellow or rarely P+ red, UV-. Chemistry: psoromic and conpsoromic acids, rarely with additional fumarprotocetraric acid.
Note: a mainly western European species ranging from Spain to Norway, found on acid soil, often in Calluna-heaths.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)