Bilimbia lobulata (Sommerf.) Hafellner & Coppins

in Veldkamp, Lichenologist, 36: 195, 2004. Basionym: Lecidea lobulata Sommerf. - Kungl. Norske Vidensk. Srifter, 2, 2: 54, 1827.
Synonyms: Bacidia sabulosa (A. Massal.) Lettau; Biatora regeliana Hepp; Bilimbia leucophaea var. perpallescens (Nyl.) A.L. Sm.; Bilimbia milliaria var. terrestris Körb.; Bilimbia regeliana (Hepp) Körb.; Bilimbia sabulosa A. Massal.; Bilimbia syncomista (Flörke) Körb.; Lecidea claudeliana Harm.; Lecidea sabuletorum var. syncomista Flörke; Lecidea subnegans Nyl.; Mycobilimbia lobulata (Sommerf.) Hafellner; Myxobilimbia lobulata (Sommerf.) Hafellner; Toninia claudeliana (Harm.) H. Olivier; Toninia lobulata (Sommerf.) Lynge; Toninia sabulosa (A. Massal.) Samp.; Toninia syncomista (Flörke) Th. Fr.; Toninia syncomista var. regeliana (Hepp) Stein
Distribution: N - Frl, Ven (Nimis 1994, Lazzarin 2000b, Nascimbene & Caniglia 2003c, Nascimbene 2005c, 2008c, Thor & Nascimbene 2007, Nascimbene & Marini 2007), TAA (Thor & Nascimbene 2007, Nascimbene 2008b), Lomb (Lazzarin 2000b), Piem (Isocrono & al. 2004, Hafellner & al. 2004), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020), Lig. C - Tosc, Marc (Nimis & Tretiach 1999), Umb (Ravera & al. 2006, 2006b), Laz (Nimis & Tretiach 2004), Abr (Nimis & Tretiach 1999, Gheza & al. 2021), Mol (Nimis & Tretiach 2004, Caporale & al. 2008), Sar. S - Camp (Ricciardi & al. 2000, Nimis & Tretiach 2004, Garofalo & al. 2010), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Ottonello & al. 1994).
Description: Thallus small-squamulose to subcrustose and warted-granular, whitish grey to grey, darker in the centre of the squamules, forming patches to 4 cm in diam. Squamules flattened, crenate to lobulate, smooth, contiguous to more or less overlapping, 0.1-0.5(-1) mm broad, forming a compact crust. Upper cortex 5-10 µm thick, overlain by an epinecral layer; algal layer with scattered groups of algae often arranged in columns; medulla white, 15-50 µm thick. Apothecia frequent, lecideine, sessile, often aggregated, 0.3-0.8(-1) mm across, with a black to brown-black, convex disc and an indistinct, soon excluded proper margin. Exciple of radiating hyphae, pale in outer part, red-brown within, K+ intensifying, N+ purple-red; epithecium pale greenish grey, K-, N+ reddish; hymenium colourless, 70-90 µm high; paraphyses mostly simple, adglutinated, 1.5-3 μm thick at mid-level, the apical cells up to 5 μm wide; subhymenium 50-75 μm high, mostly colourless in upper part, pinkish brown to dark brown in lower part; hypothecium brown-red in upper part, paler in lower part. Asci 8-spored, cylindrical-clavate, with a K/I+ blue apical dome penetrated by a narrow, K/I– apical cushion surrounded by an indistinct, apically often tapering, wall K/I– but surrounded by an I+ red-brown, K/I+ blue outer layer. Ascospores (0-)1-3-septate, hyaline, fusiform and pointed at one end, with a gelatinous coat, 12-25(-34) x 3-5(-6) µm. Photobiont chlorococcoid. Spot tests: thallus and medulla K-, C-, KC-, P-, UV-. Chemistry: thallus with zeorin.
Note: a cool-temperate to arctic-alpine, circumpolar lichen found on terricolous mosses and bare calciferous soil, most frequent in upland areas, from the Alps to the high Mediterranean mountains.
Growth form: Crustose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: rather common
Montane belt: rather rare
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

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Solar irradiation:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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P.L. Nimis CC BY-SA 4.0
TSB 38711



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (24036)
2001/12/10



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (23752)
2001/12/10



Pier Luigi Nimis CC BY-SA 4.0
TSB 41688



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (24036)



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38711



P.L. Nimis CC BY-SA 4.0
TSB 37345



P.L. Nimis CC BY-SA 4.0
TSB 37345



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (23752)
2001/12/10



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38711



P.L. Nimis CC BY-SA 4.0
TSB 38711



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38472



Pier Luigi Nimis - Department of Life Sciences, University of Trieste - CC BY-SA 4.0
TSB 37345