Brodoa atrofusca (Schaer.) Goward

Bryologist, 89: 222, 1986. Basionym: Parmelia ceratophylla var. atrofusca Schaer. - Enum. Crit. Lich. Eur.: 42, 1850.
Synonyms: Hypogymnia atrofusca (Schaer.) Räsänen; Hypogymnia intestiniformis var. atrofusca (Schaer.) Poelt; Parmelia atrofusca (Schaer.) Cromb.; Parmelia intestiniformis var. atrofusca (Schaer.) Hasselrot
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, TAA, Lomb (Nascimbene 2006), Piem (Morisi & Sereno 1995, Isocrono & al. 2004, 2006, Isocrono & Piervittori 2008), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Piervittori & al. 2001, 2004, Isocrono & al. 2008, Favero-Longo & Piervittori 2009). C - Sar.
Description: Thallus foliose, adnate, forming orbicular or irregular rosettes to 10 cm in diam. Lobes 1-2 mm broad, convex, stiff, contiguous or overlapping; center of thallus sometime with a few imbricate lobules. Upper surface pale grey to brownish, darkening or blackening especially on the lobules and towards the tips, shiny. Upper cortex paraplechtenchymatous, covered with an epinecral layer; medulla white, solid; lower cortex paraplechtenchymatous. Lower surface black, brown toward the tips, smooth to weakly and shallowly wrinkled, with scattered hapters. Apothecia rare, lecanorine, up to 10 mm diam., with a dark brown disc and a thin, entire to crenulate thalline margin. Epithecium brownish; hypothecium and hymenium colourless. Asci 8-spored, clavate, Lecanora-type. Ascospores 1-celled, hyaline, broadly ellipsoid, 7-10 x 6-7 µm. Pycnidia numerous, immersed, the visible part black to grey-brown. Conidia bifusiform, 5-7 x c. 1 µm. Photobiont chlorococcoid. Spot tests: cortex K+ yellow, C-, KC-, P-; medulla K-, C-, KC+red, P+ orange, UV-. Chemistry: cortex with atranorin and chloroatranorin; medulla with physodic and protocetraric acids.
Note: on wind-exposed surfaces of acid siliceous rocks wetted by rain near or above treeline; less bound to situations with a long snow cover than B. intestiniformis.
Growth form: Foliose, narrow lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: very rare
Oromediterranean belt: rare
Montane belt: extremely rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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P.L. Nimis CC BY-SA 4.0
TSB 13518