Calogaya lobulata (Flörke) Arup, Frödén & Søchting

in Arup & al., Nord. J. Bot., 31: 39, 2013. Basionym: Lecanora lobulata Flörke - Deutsche Lich., 3: 14, 1815.
Synonyms: Caloplaca boulyi (Zahlbr.) M. Steiner & Poelt; Caloplaca lobulata (Flörke) Hellb.; Parmelia parietina var. lobulata (Flörke) Fr.; Xanthoria boulyi Zahlbr.; Xanthoria lobulata (Flörke) B. de Lesd.; Xanthoria parietina var. lobulata (Flörke) Rabenh.; Xanthoria parietina var. turgida (Schaer.) Arnold
Distribution: N - Ven, TAA (Dalla Torre & Sarnthein 1902, Nascimbene & al. 2007b), Lomb, Piem (Isocrono & al. 2004), Lig. C - Marc (Frati & Brunialti 2006), Abr (Nimis & Tretiach 1999, Stofer 2006, Gheza & al. 2021), Mol (Caporale & al. 2008), Sar. S - Pugl (LD-1042560).
Description: Thallus at first crustose, then minutely areolate-subsquamulose to subfoliose, the areoles bright orange to grey-orange in shade-forms, smooth, sometimes somewhat shiny, minutely lobulate at margins, 0.5-1.4 mm thick, to 5 mm wide, almost completely covered with stipitate, crowded apothecia forming small pillows, and hence visible only in young specimens. Upper cortex 20-30 μm thick; medulla white; lower cortex absent. Apothecia very abundant, 1(-2) mm across, long-stipitate with hollow stalks, crowded, orbicular to irregular by mutual compression, with a flat to weakly convex, yellow-orange to brown-orange disc, and a first prominent, then often almost excluded thalline margin. Exciple 6-10 μm thick at base, 30-60 μm thick in upper part; epithecium yellowish brown, K+ red; hymenium colourless, (55-)65(-70) μm high; paraphyses distinctly branched in upper part, 1.5-2 μm thick at base, the apical cells swollen, up to 6 μm thick, inspersed with yellowish, needle-like crystals; hypothecium colourless to yellowish, underlain by a continuous algal layer. Asci 8-spored, clavate, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, ellipsoid, 10-16 x 5-7 μm, the septum 5-9 μm thick. Photobiont chlorococcoid. Spot tests: thallus and apothecia K+ red, C-, KC-, P-. Chemistry: thallus and apothecia with parietin, fallacinal, emodin, and teloschistin.
Note: a mild-temperate, subcontinental species, which in Italy is found especially on isolated almond trees, mostly on branches, in continental areas (internal Alpine and Apenninic valleys, central Sardegna). The species is included in the Italian red list of epiphytic lichens as “Vulnerable” (Nascimbene & al. 2013c).
Growth form: Crustose placodiomorph
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Subcontinental: restricted to areas with a dry-subcontinental climate (e.g. dry Alpine valleys, parts of Mediterranean Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: extremely rare


pH of the substrata:

(info)

1

2

3

4

5

Solar irradiation:

(info)

1

2

3

4

5

Aridity:

(info)

1

2

3

4

5

Eutrophication:

(info)

1

2

3

4

5

Poleotolerance:

(info)

0

1

2

3

Altitudinal distribution:

(info)

1

2

3

4

5





Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11775)
2001/12/18



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (7480)
2001/11/12



Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope "nuraghe" Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
As Caloplaca boulyi



Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
As Caloplaca boulyi



Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
As Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0144872], Syria, Muhafazat Aleppo, meadows along Euphrates river at bridge near Turkish border E of Jarablos, Populus euphratica riverside forest, arable fields, road banks, on Salix-bark. 36°49.27' N, 38°02.03' E, 400 m. Leg. L. Zedda (S-232), 3. April 1999, det. H. Sipman
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19569], Italia, Sardinia, Distr. Cagliari, Barumini, prope nuraghe Su Nuraxi, ca. 150 m, ad corticem arborum (Amygdalus communis). Leg. P.L. Nimis, C. Roux, M. Tretiach et A. Vezda, 1.09.1989. VEZDA: LICHENES SELECTI EXSICCATI NR. 2355.
as Caloplaca boulyi