Caloplaca anularis Clauzade & Poelt

Herzogia, 2: 305, 1972.
Synonyms: Caloplaca scrobiculata auct. eur. non H. Magn.
Distribution: N - Frl, Ven (Nascimbene & Caniglia 2000, 2003c, Thor & Nascimbene 2007), TAA (Nascimbene 2008b).
Description: Thallus crustose-placodioid, episubstratic, orange-yellow, to 400 μm thick, with a characteristic centrifugal growth: the thalli start dying and rapidly fall off 2-4 mm from the growing lobe-tips, leaving a 4-6 mm wide ring of 0.3-1 mm wide, contiguous, convex lobes, the free central part being often re-colonzied by younger thalli, which produces a series of concentric rings; more rarely the thalli remain rosulate, in which case they produce apothecia. Upper cortex prosoplectenchymatous, 60-140 μm thick, the upper part with many small orange crystals of anthraquinones; medulla white, up to 200 μm thick, densely filled with colourless crystals visible under polarized light; algal layer 70-150 μm thick, continuous or sometimes interrupted by cortex cones extending downwards; lower cortex poorly developed, forming a sort of hypothallus. Apothecia zeorine, 0.5-1.2 mm across, with a more less flat, orange-yellow disc and a thin, often finally excluded margin. Epithecium brownish orange, K+ red: hymenium and hypothecium colourless. Asci 8-spored, clavate, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, narrowly ellipsoid to narrowly rhomboid, (14-)15.5-17.5(-20.5) x (5-)5.7-6.5(-8) μm, the septum (1.25-)1.6-1.9(-2.25) μm thick. Pycnidia immersed, orange. Conidia 3-4.5 x 1-1.5 μm. Photobiont chlorococcoid. Spot tests: thallus and apothecia K+ red, C-, KC-, P-. Chemistry: thallus and apothecia with unidentified anthraquinones.
Note: a species of the Eurasiatic mountains, in dry-continental areas found on steeply inclined, compact limestone and dolomite, with optimum above treeline; perhaps more frequent, but undercollected in the Alps. The species differs from C. scrobiculata, of which it has been sometimes considered to be a synonym (Vondrák & Mayrhofer 2013).
Growth form: Crustose placodiomorph
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

(info)

1

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5

Solar irradiation:

(info)

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5

Aridity:

(info)

1

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3

4

5

Eutrophication:

(info)

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3

4

5

Poleotolerance:

(info)

0

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3

Altitudinal distribution:

(info)

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5





Felix Schumm - CC BY-SA 4.0
[19566], Bohemoslovacia, Slovacia, Carpates Centrales (Tatra Magna), iugum Belanské Tatry dictum, in monte Buja , alt. 1900 m, ad saxa calcarea venti exposita. Leg. E. Farkas, Z. Kyselová & A. V zda, 29.08.1985. VEZDA: LICHENES SELECTI EXSICCATI NR. 2077.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (28042)
2001/12/07



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (8217)
2001/11/12
Vezda Lich.Sel.Exs. 2077



Felix Schumm - CC BY-SA 4.0
[19566], Bohemoslovacia, Slovacia, Carpates Centrales (Tatra Magna), iugum Belanské Tatry dictum, in monte Buja , alt. 1900 m, ad saxa calcarea venti exposita. Leg. E. Farkas, Z. Kyselová & A. V zda, 29.08.1985. VEZDA: LICHENES SELECTI EXSICCATI NR. 2077.



Felix Schumm - CC BY-SA 4.0
[19566], Bohemoslovacia, Slovacia, Carpates Centrales (Tatra Magna), iugum Belanské Tatry dictum, in monte Buja , alt. 1900 m, ad saxa calcarea venti exposita. Leg. E. Farkas, Z. Kyselová & A. V zda, 29.08.1985. VEZDA: LICHENES SELECTI EXSICCATI NR. 2077.



Felix Schumm - CC BY-SA 4.0
[19566], Bohemoslovacia, Slovacia, Carpates Centrales (Tatra Magna), iugum Belanské Tatry dictum, in monte Buja , alt. 1900 m, ad saxa calcarea venti exposita. Leg. E. Farkas, Z. Kyselová & A. V zda, 29.08.1985. VEZDA: LICHENES SELECTI EXSICCATI NR. 2077.



Juri Nascimbene - CC BY-SA 4.0
Dolomiti Ampezzane, Region of Veneto, Italy
2005



Juri Nascimbene - CC BY-SA 4.0
Dolomiti Ampezzane, Region of Veneto, Italy
2005



Juri Nascimbene - CC BY-SA 4.0
Dolomiti Ampezzane, Region of Veneto, Italy
2005



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern
2007



Pier Luigi Nimis - CC BY-SA 4.0; Owner: Department of Life Sciences, University of Trieste
TSB 39011



Pier Luigi Nimis - CC BY-SA 4.0; Owner: Department of Life Sciences, University of Trieste
TSB 39011