Caloplaca nivalis (Körb.) Th. Fr.

Lichenogr. Scand., 1: 191, 1871. Basionym: Zeora nivalis Körb. - Denkschr. schles. Ber. vaterl. Kultur: 321, 1853.
Synonyms: Bacidia livida (Bagl. & Carestia) Lettau; Bilimbia livida Bagl. & Carestia; Candelariella nivalis (Körb.) Lettau; Gyalolechia nivalis (Körb.) A. Massal.; Lendemeriella nivalis (Körb.) S. Y. Kondr.; Placodium nivale (Körb.) Tuck.
Distribution: N - TAA, Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999).
Description: Thallus crustose, endosubstratic or thinly episubstratic, dirty white to grey, continuous to minutely granular, Apothecia frequent, sessile, slightly constricted at base, 0.2-0.5(-0.7) mm across, with a flat to mostly strongly convex, rusty-orange, orange-brown, olive-orange to blackish orange, pruinose disc, a granular, grey, thin, ephemeral thalline margin, and an orange to often olive-green to blackish, somehow shiny, thin, smooth proper margin. Exciple to 60 μm wide, greyish in outermost part, the pigmented parts K-, N+ red in section; proper exciple prosoplectenchymatous; epithecium orange, K+ violet-red; hymenium colourless, 80-90 μm high; paraphyses slender, 1-3 μm thick at base, not anastomosing, simple or furcate, submoniliform in upper part, slightly thickened above; hypothecium colourless, up to 110 μm high. Asci 8-spored, cylindrical-clavate, Teloschistes-type. Ascospores 1-celled to indistinctly 1-septate, hyaline, fusiform-elongate, sometimes narrower in the centre, (22-)24-38 x 3-5(-6) μm, the septum < 2 μm thick. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-; apothecia K+ red. Chemistry: thallus without lichen substances; apothecial disc with emodin, fallacinal, parietin, parietinic acid, and teloschistin; apothecial margin with the Cinereorufa-green pigment.
Note: a cool-temperate to arctic-alpine, circumpolar species found on silicicolous mosses (especially Andreaea and Grimmia). The statement of Printzen (1995) that the type of Biatora livida is Fulgensia schistidii is wrong: the type material of the former species, analyzed by M. Tretiach (pers. comm.), clearly belongs to Caloplaca nivalis.
Growth form: Crustose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

(info)

1

2

3

4

5

Solar irradiation:

(info)

1

2

3

4

5

Aridity:

(info)

1

2

3

4

5

Eutrophication:

(info)

1

2

3

4

5

Poleotolerance:

(info)

0

1

2

3

Altitudinal distribution:

(info)

1

2

3

4

5





P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (13336)
2001/11/14



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0130677], Russia, Murmansk Oblast, Kola Halbinsel, Chibiny- Gebirge, Kirowsk, Yukspor Mt. , auf Moos (Grimmia spec. ?). Leg. S. Huneck (USSR90-37), 29. June 1990, det. T. Lumbsch, 1990.


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall