Caloplaca subochracea auct.

non Caloplaca subochracea (Wedd.) Werner, Bull. Soc. bot. Fr. 102: 354 1955, nec Blastenia subochracea (Wedd.) Arup, Søchting & Frödén, Nord. J. Bot., 31: 68, 2013, nec Lecanora aurantiaca var. subochracea Wedd., Mém. Soc. Imp. Sc. Nat. Cherbourg, 17: 363, 1873.
Synonyms: Callopisma marmoratum Bagl. non auct.; Caloplaca africana sensu Clauzade; Caloplaca marmorata (Bagl.) Jatta non auct.; Caloplaca subochracea f. acrustacea Clauzade & Cl. Roux; Caloplaca subochracea f. pallida Clauzade & Cl. Roux; Caloplaca subochracea var. luteococcinea Clauzade & Cl. Roux; Chrysomma subochracea M. Choisy & Werner
Distribution: C - Tosc (Nascimbene & al. 2021), Laz (TSB 9972), Sar. S - Camp (Aprile & al. (2003b), Pugl (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Nimis & al. 1994, Ottonello & Salone 1994, Ottonello & al. 1994, Grillo & al. 2002, Grillo & Caniglia 2004, Caniglia & Grillo 2005, 2006, Genco & al. 2007).
Description: Thallus crustose, very variable, from endosubstratic and poorly evident to episubstratic and finely rimose-areolate, the colour varying from pure white to pale or bright yellow-orange, often delimited by a thin, black prothalline line. Apothecia biatorine, at first immersed, then sessile, rounded, 0.2-1.3 mm across, with a rusty red to dark orange-red, at first flat, then convex, epruinose disc, and a somewhat paler, smooth, at first well-visible, finally often excluded proper margin. Epithecium brownish orange, K+ red; hymenium colourless, 40-60(-80) µm high; paraphyses simple or forked in upper part, c. 1 µm thick at base, the apical cells 2-3 µm wide; hypothecium colourless. Asci 8-spored, clavate, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, ellipsoid, (9-)10-13(-14) x (3-)5-6(-10) µm, the septum up to 5 µm thick. Photobiont chlorococcoid. Spot tests: thallus K- or K+ red; C-, KC-, P-; Apothecia K+ red. Chemistry: apothecia (and sometimes thallus) with unidentified anthraquinones.
Note: a Mediterranean species found on compact limestone, locally very abundant in coastal situations, extremely rare far from the coast. The colour of the thallus, based on which several infraspecific taxa were distinguished (see e.g. Roux & coll. 2014), is variable depending on exposure to sunlight, and intermediate forms are frequent. There are open issues with the epithet of this lichen (see Roux & coll. 2014). I have seen the type of Callopisma marmoratum Bagl. (MOD-TSB), which clearly belongs to this lichen and not to that which is usually called “Caloplaca marmorata” by most authors.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Taxon bound to maritime-coastal situations

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: rather common
Dry mediterranean belt: rather rare


pH of the substrata:

(info)

1

2

3

4

5

Solar irradiation:

(info)

1

2

3

4

5

Aridity:

(info)

1

2

3

4

5

Eutrophication:

(info)

1

2

3

4

5

Poleotolerance:

(info)

0

1

2

3

Altitudinal distribution:

(info)

1

2

3

4

5





P.L. Nimis CC BY-SA 4.0
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari.
Herbarium: Holotype of Callopisma marmoratum Bagl (MOD)
09.06.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari.
Herbarium: Holotype of Callopisma marmoratum Bagl (MOD)
09.06.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari.
Herbarium: Holotype of Callopisma marmoratum Bagl (MOD)
09.06.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari.
Herbarium: Isotype of Callopisma marmoratum Bagl. (MOD)
09.06.2016



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Sicily, Trapani, Erice, Historical Center


Pier Luigi Nimis
Italy, Sicily, Trapani, Erice, old city walls near the Castle
12/2016



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sicily, Trapani, Erice, castle
12/2016



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sicily, Trapani, Erice, castle
12/2016



Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710.
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710.
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710.
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972.
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972.
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972.
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea


Department of Life Sciences, University of Trieste
Herbarium: TSB (11674)
2001/11/14



P.L. Nimis CC BY-SA 04
TSB 36720



P.L. Nimis CC BY-SA 04
TSB 36720