Gyalolechia marmorata (Bagl.) Nimis & Arup

in Arup & al., Borziana, 4: 45, 2023. Basionym: Callopisma marmoratum Bagl. - N. Giorn. Bot. Ital., 11: 84, 1879
Synonyms: Caloplaca marmorata (Bagl.) Jatta non auct.; Caloplaca subochracea auct.; Caloplaca subochracea f. acrustacea Clauzade & Cl. Roux; Caloplaca subochracea f. pallida Clauzade & Cl. Roux; Caloplaca subochracea var. luteococcinea Clauzade & Cl. Roux
Distribution: C - Tosc (Nascimbene & al. 2021), Laz (TSB 9972), Sar (Arup & al. 2023). S - Camp (Aprile & al. (2003b), Pugl (Nimis & Tretiach 1999, Arup & al. 2023), Cal (Puntillo 1996), Si (Nimis & al. 1994, Ottonello & Salone 1994, Ottonello & al. 1994, Grillo & al. 2002, Grillo & Caniglia 2004, Caniglia & Grillo 2005, 2006, Genco & al. 2007, Arup & al. 2023).
Description: Thallus crustose, very variable, from endosubstratic and poorly evident to episubstratic and finely rimose-areolate, white to pale yellow, often delimited by a thin, black prothalline line. Apothecia biatorine, at first immersed, then sessile, rounded, 0.2-1.3 mm across, with a rusty red to dark orange-red, at first flat, then convex, epruinose disc, and a somewhat paler, smooth, at first well-visible, finally often excluded proper margin. Epithecium brownish orange, K+ red; hymenium colourless, 40-60(-80) µm high; paraphyses simple or forked in upper part, c. 1 µm thick at base, the apical cells (2-)3-4.5 µm wide; hypothecium colourless. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, ellipsoid, (9-)10-13(-14) x (3-)5-6(-10) µm, the equatorial thickening (“septum”) up to 5 µm. Photobiont chlorococcoid. Spot tests: thallus K- or K+ faintly red; C-, KC-, P-; Apothecia K+ red. Chemistry: thallus without lichen substances or with low amounts of fragilin; apothecia with parietin and emodin (major), and citreorosein, emodinal, fallacinal and parietinic acid (minor).
Note: a Mediterranean species found on compact limestone, locally very abundant in coastal situations, rare far from the coast (Arup & al.2023).
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Taxon bound to maritime-coastal situations

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: rather common
Dry mediterranean belt: rather rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Michel Bertrand
France


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016



P.L. Nimis CC BY-SA 04
TSB 36720



P.L. Nimis
holotype of Callopisma marmoratum Bagl. 1879



P.L. Nimis CC BY-SA 4.0
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis CC BY-SA 04
Herbarium: TSB 36720



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)