Caloplaca ulcerosa Coppins & P. James

Lichenologist, 11: 139, 1979.
Synonyms: Coppinsiella ulcerosa (Coppins & P. James) S.Y. Kondr. & L. Lőkös
Distribution: N - VG, Frl (Nimis & al. 1996), Ven (vidi!), Lomb (vidi!), Piem (TSB 33520), Emil (Nimis & al. 1996, Fariselli & al. 2020), Lig (TSB 21798). C - Tosc (Tretiach & Nimis 1994, Frati & al. 2008, Benesperi 2011), Marc (Nimis & Tretiach 1999), Umb (Ravera & Di Toma 2003, Ravera & al. 2006), Laz, Abr (Nimis & Tretiach 1999), Mol (Nimis & Tretiach 1999, Caporale & al. 2008), Sar. S - Camp (Nimis & Tretiach 2004, Garofalo & al. 2010), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999, Paoli & al. 2006), Cal, Si (Ottonello & Salone 1994, Grillo & Cataldo 2008).
Description: Thallus crustose, thin, continuous, grey-white, with yellowish green, excavated, spindle-shaped to round, greenish soralia bearing coarsely farinose soredia. Apothecia not uncommon, zeorine, up to 0.5(-0.7) mm across, with a bright orange disc, a yellow-orange, finally often excluded proper margin and sometimes a poorly developed, finally disappearing, grey-white thalline margin. Epithecium orange, K+ red; hymenium colourless; paraphyses simple or branched in upper part, 1-3 μm thick, the upper cells to 6 μm wide; hypothecium colourless. Asci 8-spored, clavate, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, broadly ellipsoid, 8-12 x 4-8 µm, the septum 3-6 µm thick (1/3-1/2 of spore length). Photobiont chlorococcoid. Spot tests: thallus and soralia K-, C-, KC-, P-: apothecia K+ red. Chemistry: unidentified anthraquinones in the apothecia.
Note: a warm-temperate, rather misunderstood but well-distinct species, especially common on the rough bark of isolated trunks of Ulmus and Populus, often along white roads and usually not far from the coast (Vondrák & al. 2009b). The species will be soon transferred into a new genus (Vondrák in litt.).
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: rather rare
Padanian area: extremely rare
Humid submediterranean belt: rare
Humid mediterranean belt: very rare
Dry mediterranean belt: extremely rare


pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

(info)

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Altitudinal distribution:

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Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (2998)
2001/11/19



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29815)
2008.02.25



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29998)
2008.02.25



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (25288)
2008.02.25



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (25288)
2008.02.25



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (25288)
2008.02.25



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (21749)
2008.03.03



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (21749)
2008.03.03



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29815)
2001/12/07



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95



Felix Schumm – CC BY-SA 4.0
[19407], USA, South Dakota, Grant Co., Blue Cloud Benedictine Abbey ca. 1.6 km E of Marvin, open pastureland, 45°15'05'' N, 95°53'28'' W, 467 m, on deciduous trees. Leg. M.K. Advaita (No. 4915), 18. May 2005, det. C. Wetmore. TELOSCHISTACEAE EXS. 95