Cladonia cervicornis (Ach.) Flot.

Jahrb. schles. Ges. vaterl. Kultur, 27: 105, 1849. Basionym: Lichen cervicornis Ach. - Lichenogr. Suec. Prodr.: 198, 1799.
Synonyms: Cladonia cervicornis f. epiphylla (Rabenh.) Clauzade & Cl. Roux; Cladonia cervicornis f. sobolifera (Nyl.) Delise; Cladonia sobolifera (Delise) Nyl.; Cladonia verticillata var. cervicornis (Ach.) Flörke
Distribution: N - Ven, TAA (Nascimbene & al. 2022), Lomb (Rivellini & Valcuvia 1996, Rossi & al. 1998, Gheza 2015, 2018, 2019, 2019b, Gheza & al. 2022), Piem (Isocrono & al. 2004, Gheza 2015, 2018), VA (Piervittori & Isocrono 1997, 1999, Valcuvia 2000), Lig (Gheza & al. 2020). C - Tosc (Pišút 1997), Laz, Sar (Nöske 2000, Zedda & al. 2010, Cogoni & al. 2011, Neuwirth 2018). S - Camp, Pugl (Durini & Medagli 2004, Gianfreda & Matino 2020), Bas (Potenza 2006, Potenza & Fascetti 2012), Cal (Puntillo 1996, Puntillo & Puntillo 2004), Si (Ottonello & Romano 1997, Merlo 2004, Caniglia & al. 2005, Brackel 2008b, Ottonello & al. 2011).
Description: Primary thallus squamulose, persistent, usually forming 5-10 cm wide, dense pillows or mats, the squamules large, to 2.5 cm long, 1-4 mm wide, contiguous, ascending, grey-green above, brownish or bluish grey and very finely arachnoid beneath, with entire to indented margins. Podetia rather rare, greyish to brownish green, sometimes slightly melanotic at base, funnel-shaped, 7-20(-30) mm tall, flaring from the base, often proliferating from the centre with one to several additional cups originating from a single cup (forming 1-4 superimposed tiers), more rarely proliferating from the margins, with a continuous to areolate cortex, sometimes sparingly squamulose at the base. Apothecia rare, dark brown, convex, located on the margins of cups. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed in the margins of cups, subglobose to ovoid, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P+ red, UV-. Chemistry: fumarprotocetraric acid complex.
Note: a temperate to southern boreal-montane lichen found on mineral siliceous soil in open grasslands and garrigues, most frequent in Tyrrhenian Italy. See also note on C. verticillata.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: rather common
Dry mediterranean belt: extremely rare

pH of the substrata:

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Solar irradiation:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C3/Cladonia_cervicornis.htm
France, session Cladonia 2013 - Fontainebleau



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (38339)
2008.02.25



Felix Schumm – CC BY-SA 4.0
[9128], Spanien, Kanarische Inseln, La Gomera, beim Mirador de Alojera (Piedras Hincadas) nördlich von Arure im Lorbeer-Erica Wald, 28°08.995' N, 17°18.483’ W, 1000 m. Leg. Schumm 10.02.2002, det Ahti 2011.



Felix Schumm – CC BY-SA 4.0
[9128], Spanien, Kanarische Inseln, La Gomera, beim Mirador de Alojera (Piedras Hincadas) nördlich von Arure im Lorbeer-Erica Wald, 28°08.995' N, 17°18.483’ W, 1000 m. Leg. Schumm 10.02.2002, det Ahti 2011.



Felix Schumm – CC BY-SA 4.0
[9128], Spanien, Kanarische Inseln, La Gomera, beim Mirador de Alojera (Piedras Hincadas) nördlich von Arure im Lorbeer-Erica Wald, 28°08.995' N, 17°18.483’ W, 1000 m. Leg. Schumm 10.02.2002, det Ahti 2011.



Felix Schumm – CC BY-SA 4.0
[9128], Spanien, Kanarische Inseln, La Gomera, beim Mirador de Alojera (Piedras Hincadas) nördlich von Arure im Lorbeer-Erica Wald, 28°08.995' N, 17°18.483’ W, 1000 m. Leg. Schumm 10.02.2002, det Ahti 2011.,
pr: protocetraric acid, fpr: fumarprotocetraric acid


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (13288)
2003/02/15
prymary thallus


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C3/Cladonia_cervicornis.htm
France, session Cladonia 2013 - Fontainebleau



P.L. Nimis CC BY-SA 4.0
TSB 9937


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C3/Cladonia_cervicornis.htm
France, session Cladonia 2013 - Fontainebleau