Cladonia fimbriata (L.) Fr.

Lichenogr. Eur. Ref.: 222, 1831. Basionym: Lichen fimbriatus L. - Sp. Pl., 2: 1152, 1753.
Synonyms: Cladonia fimbriata var. chordalis auct.; Cladonia major (K.G. Hagen) Sandst.; Cladonia minor K.G. Hagen
Distribution: N - VG, Frl (Tretiach & Molaro 2007, Brackel 2013), Ven (Nascimbene & Caniglia 1997, 2002c, 2003c, Caniglia & al. 1999, Nascimbene 2005c, 2008, 2011, Nascimbene & al. 2005b, 2006, 2006c, 2006e, 2007, 2009c, 2010b, 2013b, 2015, Nascimbene & Marini 2007), TAA (Caniglia & al. 2002, Nascimbene & Caniglia 2002c, Nascimbene 2003, 2005b, 2006b, 2006c, 2008b, 2008c, 2013, 2014, Nascimbene & al. 2006e, 2007c, 2008c, 2014, Lang 2009, Nimis & al. 2015), Lomb (Rivellini 1994, Rivellini & Valcuvia 1996, Zocchi & al. 1997, Roella 1999, De Vita & Valcuvia 2004, Dalle Vedove & al. 2004, Anderi & al. 2005, Nascimbene & al. 2006e, Stofer 2006, Furlanetto 2010, Gheza & al. 2015, Gheza 2015), Piem (Caniglia & al. 1992, Morisi & Sereno 1995, Isocrono & al. 2003, 2004, Giordani & Malaspina 2016), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Valcuvia 2000, Piervittori & al. 2001, Matteucci & al. 2008, Furlanetto 2010), Emil (Nimis & al. 1996, Dalle Vedove & al. 2002, Tretiach & al. 2008, Benesperi 2009, Fariselli & al. 2020), Lig (Brunialti & al. 1999, Giordani 2006, Giordani & Incerti 2008, Gheza & al. 2020). C - Tosc (Tretiach & Nimis 1994, Loppi & Putortì 1995b, Loppi & Nascimbene 1998, 2010, Putortì & Loppi 1999b, Benesperi 2000a, 2006, Paoli & Loppi 2001, Loppi & al. 2002c, Lorenzini & al. 2003, Loppi & Frati 2006, Benesperi & al. 2007, Lastrucci & al. 2009, Pasquinelli & al. 2009, Pasquinelli & Puccini 2010, Benesperi 2011, Brackel 2015, Gheza & al. 2020), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Panfili 2000b, 2007, Ravera & al. 2006), Laz (Bartoli & al. 1997, Ravera & al. 1999, Ravera 2002, Massari & Ravera 2002, Ruisi & al. 2005, Munzi & al. 2007, Genovesi & al. 2011, Zucconi & al. 2013), Abr (Nimis & Tretiach 1999, Caporale & al. 2016, Gheza & al. 2021), Mol (Garofalo & al. 1999, Caporale & al. 2008, Genovesi & Ravera 2014, Paoli & al. 2015, Brackel 2020), Sar (Zedda 1995, 2002, Zedda & Sipman 2001, Rizzi & al. 2011). S - Camp (Garofalo & al. 1999, 2010, Ricciardi & al. 2000, Aprile & al. 2003, 2003b, Nimis & Tretiach 2004, Brunialti & al. 2010, 2013, Nascimbene & al. 2010b, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Garofalo & al. 1999, Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1995, 1996, Puntillo & Puntillo 2004, Incerti & Nimis 2006), Si (Nimis & al. 1994, Ottonello & Salone 1994, Ottonello & al. 1994, 2011, Grillo & al. 1996, Ottonello 1996, Ottonello & Romano 1997, Grillo 1998, Merlo 2004, Grillo & Caniglia 2004, 2006, Falco Scampatelli 2005, Stofer 2006, Liistro & Cataldo 2011).
Description: Primary thallus squamulose, persistent or ephemeral, the squamules middle-sized to rather large, 2-4(-5) mm long, 1-3 mm wide, elongate and incised, grey green above, white beneath, sometimes sorediate. Podetia goblet-shaped, hollow inside, green to grey-green, mostly ecorticate and esquamulose, 0.5-3 cm tall, with small, regular, rarely marginally proliferating cups (c. 0.5 cm diam.) and long stalks which are 1-2 mm thick at base, farinose-sorediate throughout, the soredia 30-50 μm in diam. Apothecia rare, dark brown, convex, sometimes aggregated into 1-5 mm wide clusters. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia brown, mainly found along the margins of cups. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P+ red, UV-. Chemistry: fumarprotocetraric acid, sometimes with additional quaesitic acid.
Note: a widespread temperate to arctic-alpine, holarctic species found on rotten wood, soil, at the base of trunks, with a wide ecological range and a correspondingly wide altitudinal range.
Growth form: Fruticose

Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: rather rare
Oromediterranean belt: rare
Montane belt: very common
Submediterranean belt: common
Padanian area: very rare
Humid submediterranean belt: very common
Humid mediterranean belt: common
Dry mediterranean belt: extremely rare


pH of the substrata:

(info)

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Solar irradiation:

(info)

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Aridity:

(info)

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Eutrophication:

(info)

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Poleotolerance:

(info)

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Altitudinal distribution:

(info)

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Tomi Trilar; Owner: Slovenian Museum of Natural History
Switzerland, Valais, Leuk, Forêt de Finges (Pfynwald)
08.06.2007



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10747)
2001/12/04


Andres Saag; Owner: University of Tartu


Triin Aimla; Owner: University of Tartu



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Veneto, BL, Comune di Sappada, pendici boscose del Monte Peralba.
2008.27.08



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Veneto, BL, Comune di Sappada, pendici boscose del Monte Peralba.
2008.27.08



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Veneto, BL, Comune di Sappada, pendici boscose del Monte Peralba.
2008.27.08



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Veneto, BL, Comune di Sappada, pendici boscose del Monte Peralba.
2008.27.08



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Veneto, BL, Comune di Sappada, pendici boscose del Monte Peralba.
2008.27.08



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Veneto, BL, Comune di Sappada, pendici boscose del Monte Peralba.
2008.27.08



Tomi Trilar; Owner: Slovenian Museum of Natural History
Switzerland, Valais, Leuk, Forêt de Finges (Pfynwald)
08.06.2007



Tomi Trilar; Owner: Slovenian Museum of Natural History
Switzerland, Valais, Leuk, Forêt de Finges (Pfynwald)
08.06.2007



Tomi Trilar; Owner: Slovenian Museum of Natural History
Switzerland, Valais, Leuk, Forêt de Finges (Pfynwald)
08.06.2007



Tomi Trilar; Owner: Slovenian Museum of Natural History
Switzerland, Valais, Leuk, Forêt de Finges (Pfynwald)
08.06.2007


Antonio Valero; Owner: Antonio Valero
Spain
2010


Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, San Rossore
2010


Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Area Naturale Protetta Poggio Ripaghera - S. Brigida - Valle dell’Inferno
2010


Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010


Giancarlo Medici; Owner: Giancarlo Medici
Italia, Alpi.
Herbarium: Forum Natura Mediterraneo
2010


Giancarlo Medici; Owner: Giancarlo Medici
Italia, Alpi.
Herbarium: Forum Natura Mediterraneo
2010


Nere Sorrentino
Spain, Canary Islands


Nere Sorrentino
Spain, Canary Islands


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Varese, Somma Lombardo, 228 m
05/11/2020