Enchylium bachmanianum (Fink) Otálora, P.M. Jørg. & Wedin

Fungal Divers. (online version), 64: 286, 2013. Basionym: Collemodes bachmanianum Fink - Mycologia, 10: 236, 1918.
Synonyms: Collema bachmanianum (Fink) Degel.; Collema bachmanianum var. millegranum Degel.
Distribution: N - Lomb.
Description: Thallus foliose, homoiomerous, gelatinous when wet, 2-6 cm wide, loosely adnate, dark olive-green to black; lobes 2-4(-6) mm wide, radiating, concave to flat, 200-700 µm thick (when wet), sometimes with many isidia-like warts or lobules. Apothecia lecanorine, sessile and constricted at base, 1.5-3(-4) mm across, with a concave to flat, reddish brown disc, and a coarsely crenulate to lobulate, usually persisting thalline margin. Proper exciple thin, euthyplectenchymatous; epithecium brownish; hymenium colourless, 100-135 µm high, I+ blue; hypothecium colourless. paraphyses simple or sparingly branched, with clavate to subglobose tips; hypothecium colourless. Asci 8-spored, clavate, the apex thick, with a I+ blue apical dome and a I+ blue annulus projecting downwardly. Ascospores submuriform, mostly with 3 transverse septa and 1(-2) longitudinal septa, at first hyaline, then pale yellow or brown, ellipsoid with usually pointed ends, (20-)26-32(-36) x 8.5-15 µm. Photobiont cyanobacterial (Nostoc, the cells arranged in long chains). Spot tests: all negative. Chemistry: without lichen substances.
Note: a mainly arctic-alpine, circumpolar species found on calciferous or base-rich siliceous soil near or above treeline; restricted to the Alps in Italy. Italian samples belong to var. millegranum.
Growth form: Foliose, broad lobed
Substrata: soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: extremely rare
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

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Poleotolerance:

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Altitudinal distribution:

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Curtis Randall Björk – CC BY-SA 4.0
Clearwater Valley, BC, First Canyon Uplands Date: 2011-05-05: On cliffs of pillow basalt on south-facing slope, dry, lightly forested



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU - Lichenes Alpium (280)
2003/04/28
apothecium



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU - Lichenes Alpium (280)
2003/04/28


Degelius G. 1954. The lichen genus Collema in Europe: Morphology, Taxonomy, Ecology. Symbolae Bot. Upsal. 13, 2: 1-499.