Synonyms: Heppia gigantea Egea & Llimona; Heppia paulina Marton; Psora lobatoplicata B. de Lesd.
Distribution: N - Lig.
Description: Thallus squamulose, heteromerous, 340-400 µm thick, ochraceous brown, with pale punctiform depressions in a reticulate pattern. Squamules subpeltate, 2.5-7 mm wide, at first flat, then concave, contiguous, with rounded ends and a down-turned edges, attached by a mat of 10-14 µm thick, colourless rhizohyphae, forming 1.4-2.7(-5) cm wide, convex rosettes. Upper cortex paraplectenchymatous, (25-)50-75 µm thick, covered with a 10-15 µm thick epinecral layer; medulla poorly developed, of anticlinally arranged hyphae; lower cortex absent or restricted to the margins. Apothecia without a thalline margin, semi-immersed in the squamules, 0.5-1.5(-2) mm across, with a dark reddish brown, concave disc, and a thin to indistinct proper margin. Proper exciple very thin, colourless; epithecium brownish, K-; hymenium colourless, 120-180 µm high, I+ blue slowly turning reddish; paraphyses simple, lax, 4-6 µm thick at base, the apical cells 8-10 µm wide; hypothecium colourless, I+ blue. Asci 8-spored, thin-walled, narrowly clavate to subcylindrical, prototunicate. Ascospores 1-celled, hyaline, ellipsoid, thin-walled, (18-)20-29(-31) x (7-)9-12 µm. Pycnidia dark, immersed. Conidia fusiform. Photobiont cyanobacterial, Scytonema-like. Spot tests: all negative. Chemistry: without lichen substances.
Note: the Italian record is the northernmost known for this Mediterranean-Macaronesian lichen growing on clay soil in dry, open grasslands. According to Timdal (in litt.) Psora lobatoplicata is a likely synonym of this species.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris
[ABL77805], Brazil, Mato Grosso do Sul, Bodoquena, Dente de Cão
summit,in Atlantic rain forest on exposed limestone. 20°47’´05'' S,
56°45’03'' W, 450 m. Leg. A. Aptroot (no 77805), 7.11.2018, det. A.