Distribution: C - Tosc (TSB 35171). S - Pugl (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Caniglia & Grillo 2001, 2005, 2006, Grillo & Caniglia 2004, 2005, Cataldo & Minissale 2013, 2015).
Description: Thallus squamulose, brownish under a thick white layer of clustered crystals (pruina) which tends to accumulate in areolate patches, giving the surface a reticulate appearance. Squamules (2-)3-7 mm broad, 300-400 µm thick, elongate, at first flat, then concave, contiguous, with rounded ends and up-turned, undulate edges, attached by a mat of rhizohyphae. Upper cortex paraplectenchymatous, 35-75 µm thick, overlain with a 15-25 v thick epinecral layer; medulla 60.100 µm thick, paraplectenchymatout; lower cortex 25-30 µm thick, paraplectenchymatous, hardly differentiated from the medulla. Apothecia frequent, without a thalline margin, semi-immersed in the squamules, 0.5-1.5(-2.5) mm across, with a reddish brown, at first concave, then flat disc. Proper exciple poorly developed: epithecium orange-brown, K-; hymenium colourless, 150-200 µm high; paraphyses mostly simple, lax, distinctly thickened above; hypothecium colourless. Asci 8-spored, cylindrical to obovoid, prototunicate, thin-walled, the wall I+ blue. Ascospores 1-celled, hyaline, ellipsoid, thin-walled, 16-26(-29) x (6-)8-12(-13) µm. Pycnidia dark, immmersed. Conidia bacilliform. Photobiont cyanobacterial (Scytonema-like). Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: on clay or sandy-clay soil, restricted to very dry grasslands in Mediterranean Italy.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris