Distribution: N - TAA, Piem (Isocrono & al. 2004), Lig.
Description: Thallus squamulose to subcrustose, brownish, sometimes with a grey tinge, dull, thin. Squamules <1.5 mm broad and long, usually contiguous and forming a crust, flattened to slightly convex, smooth, irregular in shape, the margins entire to usually crenate-lobulate, not up-turned, surrounded by a dark hypothallus, the marginal squamules often somehow elongate. Lower surface dark, attached by 4-5 µm thick, dark rhizohyphae. Upper and lower cortex paraplectenchymatous; medulla not differentiated. Perithecia frequent, developing on a dark hypothallus among the squamules, globose, projecting, with an involucrellum extending all around the perithecium, 0.2-0.35 mm across. Exciple black and carbonaceous throughout; paraphyses absent, substituted by periphyses. Asci 8-spored, clavate, the wall I-. Ascospores 1-celled, hyaline, narrowly ovoid to club-shaped, thin-walled, (15-)17-21(-23) x (7.5-)8-10(-11) µm, biseriately arranged in the asci. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P- UV-. Chemistry: without lichen substances.
Note: on calciferous, humus-rich soil in alpine grasslands, probably more widespread in the Alps.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris