Melanohalea elegantula (Zahlbr.) O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch

Mycol. Res., 108: 882, 2004. Basionym: Parmelia olivacea subsp. aspidota var. elegantula Zahlbr. - Verh. Ver. Nat. Heilk. Pressburg, 8: 39, 1894.
Synonyms: Collema exasperatum Ach.; Melanelia elegantula (Zahlbr.) Essl.; Melanelia incolorata (Parrique) Essl.; Parmelia elegantula (Zahlbr.) Szatala; Parmelia exasperatula var. elegantula (Zahlbr.) Zahlbr.; Parmelia incolorata (Parrique) Lettau; Parmelia jacquesii Werner
Distribution: N - VG (Castello 1996, Castello & Skert 2005), Frl (Tretiach & Molaro 2007), Ven (Nascimbene & Caniglia 1997, Caniglia & al. 1999, Nascimbene 2005c, Nascimbene & al. 2009c, Brackel 2013), TAA (Nascimbene 2003, Nascimbene & al. 2007b), Lomb (Rivellini 1994, Arosio & Rinaldi 1995, Zocchi & al. 1997, Roella 1999, Arosio & al. 2003, Stofer 2006, Brackel 2010, Furlanetto 2010, Gheza 2019b), Piem (Morisi & Sereno 1995, Arosio & al. 1998, Isocrono & Falletti 1999, Piervittori 2003, Isocrono & al. 2004, 2007, Isocrono & Piervittori 2008, Furlanetto 2010, Matteucci & al. 2010, Giordani & Malaspina 2016), VA (Piervittori & Maffei 1996, Piervittori & Isocrono 1999, Valcuvia & al. 2000b, Matteucci & al. 2008, 2008c), Emil (Bassi 1995, Tretiach & al. 2008, Benesperi 2009, Gerdol & al. 2014, Fariselli & al. 2020), Lig (Brunialti & Giordani 2003). C - Tosc (Loppi & al. 1994, 1997, 1997b, 1998b, 2002, 2003, 2004c, 2006, Loppi & Putortì 1995b, Monaci & al. 1997, Loppi & Nascimbene 1998, 2010, Tretiach & Ganis 1999, Benesperi & al. 2007, Brunialti & Frati 2010, Benesperi 2011, Paoli & al. 2012, Brackel 2015), Marc (Gasparo & al. 1989, Nimis & Tretiach 1999, Brackel 2015), Umb (Ravera 1998, Ravera & al. 2006), Laz (Ravera 2001, 2002, 2008b, Massari & Ravera 2002, Ruisi & al. 2005, Munzi & al. 2007, Ravera & Genovesi 2008, Zucconi & al. 2013, Brackel 2015), Abr (Nimis & Tretiach 1999, Brackel 2015, Caporale & al. 2016, Corona & al. 2016, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Ravera, Caporale & al. 2008, Genovesi & Ravera 2014, Brackel 2020, Caporale & Ravera 2020), Sar (Zedda 1995, 2002, Loi & al. 2000, Rizzi & al. 2011, Brackel & Berger 2019). S - Camp (Aprile & al. 2002, 2003, Nimis & Tretiach 2004, Brunialti & al. 2010, 2013, Nascimbene & al. 2010b, Garofalo & al. 2010, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999, Brackel 2011), Bas (Nimis & Tretiach 1999, Fascetti & al. 2005, Potenza 2006, Potenza & al. 2010, Brackel 2011), Cal (Puntillo 1995, 1996, Incerti & Nimis 2006, Stofer 2006, Brackel & Puntillo 2016), Si (Grillo & al. 1996, Grillo 1998, Grillo & Caniglia 2004, 2006, Falco Scampatelli 2005).
Description: Thallus foliose, heteromerous, dorsiventral, forming 4-7 cm wide rosettes, isidiate. Lobes 2-7 mm wide, contiguous to imbricate, adpressed but sometimes slightly ascending at margins, brown, usually dull, but sometimes shiny at lobe tips, epruinose, with abundant, cylindrical, simple to coralloid, 0.05-0.18 mm thick, not hollow isidia. Lower surface pale brown, slightly darker at margin, with mostly simple rhizines. Upper cortex paraplectenchymatous, with a non-pored epicortex, the cell walls containing isolichenan; medulla white; lower cortex paraplectenchymatous. Apothecia very rare, lecanorine, with a brown disc and an isidiate thalline margin. Epithecium brown; hymenium and hypothecium colourless. Asci 8-spored, clavate, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid, 8-12 x 4-7 µm. Pycnidia rare, black, laminal. Conidia slightly dumbbell-shaped, c. 7 x 1 µm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, N-, UV-. Chemistry: without lichen substances.
Note: a mild-temperate lichen found on old trees (e.g. oaks, Castanea), more rarely on siliceous rocks (Roux & coll. 2014), with optimum in the montane belt; common in the Apennines, rare in northern Italy, and absent from urban areas.
Growth form: Foliose, broad lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather common
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 04
TSB 10984



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31232)
2001/12/21
tip of lobes



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28


Antonio Valero; Owner: Antonio Valero
Spain
2010


Pier Luigi Nimis - CC BY-SA 4.0
Herbarium: TSB 17796



Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.



Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.



Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.



Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.



Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118



Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118



Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118



Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335



Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335



Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335



P.L. Nimis CC BY-SA 4.0
TSB 83637



P.L. Nimis CC BY-SA 4.0
TSB 83637



P.L. Nimis CC BY-SA 4.0
TSB 10984


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/