Brodoa atrofusca (Schaer.) Goward

Bryologist, 89: 222, 1986. Basionym: Parmelia ceratophylla var. atrofusca Schaer. - Enum. Crit. Lich. Eur.: 42, 1850.
Synonyms: Hypogymnia atrofusca (Schaer.) Räsänen; Hypogymnia intestiniformis var. atrofusca (Schaer.) Poelt; Parmelia atrofusca (Schaer.) Cromb.; Parmelia intestiniformis var. atrofusca (Schaer.) Hasselrot
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, TAA (Nascimbene & al. 2022), Lomb (Nascimbene 2006, Gheza 2017), Piem (Morisi & Sereno 1995, Isocrono & al. 2004, 2006, Isocrono & Piervittori 2008), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Piervittori & al. 2001, 2004, Isocrono & al. 2008, Favero-Longo & Piervittori 2009). C - Sar.
Description: Thallus foliose, heteromerous, dorsiventral, adnate, forming up to 5 (rarely more) cm wide, orbicular or irregular rosettes. Lobes 1-2 mm broad, convex, stiff, contiguous or overlapping; center of thallus sometimes with a few imbricate lobules. Upper surface pale whitish, pale grey to brownish, darkening or blackening especially on the lobules and towards the tips, somehow glossy. Lower surface black, brown toward the tips, smooth to weakly and shallowly wrinkled, erhizinate, with scattered hapters. Upper cortex paraplectenchymatous, the cell walls with Cetraria-type lichenan, with a non-pored epicortex; medulla white; lower cortex paraplectenchymatous, without an epicortex. Apothecia rare, lecanorine, up to 10 mm across, with a dark brown disc and a thin, entire to crenulate thalline margin. Epithecium brownish; hypothecium and hymenium colourless; paraphyses mostly simple. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, broadly ellipsoid, (7-)9-11(-12) x 6-7(-8) µm. Pycnidia numerous, immersed, the visible part black to grey-brown. Conidia bifusiform, 5-7 x c. 1 µm. Photobiont chlorococcoid. Spot tests: upper cortex K+ yellow, C-, KC-, P-; medulla K-, C-, KC+ red, P+ orange, UV-. Chemistry: upper cortex with atranorin and chloroatranorin; medulla with physodic and protocetraric acids.
Note: on wind-exposed surfaces of acid siliceous rocks wetted by rain near or above treeline; less bound to situations with a long snow cover than B. intestiniformis.
Growth form: Foliose, narrow lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: very rare
Oromediterranean belt: rare
Montane belt: extremely rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model
Herbarium samples

P.L. Nimis CC BY-SA 4.0
TSB 13518