Kuettlingeria furax (Egea & Llimona) I.V. Frolov, Vondrák & Arup

in Frolov & al., J. Syst. Evol. 59, 3: 468, 2020. Basionym: Caloplaca furax Egea & Llimona -
Synonyms: Caloplaca furax Egea & Llimona; Pisutiella furax (Egea et Llimona) S. Y. Kondr., L. Lőkös & E. Farkas
Distribution: C - Sar (Nimis 1993). S - Bas (Nimis & Tretiach 1999), Si (Nimis & al. 1996b).
Description: Thallus crustose-subsquamulose, dark ash grey to lead-grey, dull, episubstratic, 135-175 μm thick, developing upon a black hypothallus on the thalli of aspicilioid lichens, forming 2-6 mm wide, often confluent patches with a sublobulate margin, tending to die in central parts and becoming ring-like. Cortex 27-30 μm thick, the upper part violaceous grey, covered with a hyaline, 14-20 μm thick epinecral layer. Apothecia zeorine/lecanorine, 0.4-0.8 mm across, usually numerous, at first immersed and clearly provided with a thalline margin, then subsessile and with a rather thick, persistent, bright orange proper margin, the disc at first concave, then flat to slightly convex, orange to orange-brown. Proper exciple orange, corticate in outer part, K+ red; epithecium orange-brown, K+ red, C+ purple, hymenium colourless, 80-90 μm high; paraphyses sparingly branched in upper part, the uppermost cells slightly swollen, 2-3 µm wide; hypothecium colourless. Asci 8-spored, clavate, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, broadly ellipsoid to subglobose, often swollen at septum, (11.5-)12-15(-16) x 7-10 μm, the septum 4-5.5 μm thick. Pycnidia immersed in small thalline warts. Conidia broadly fusiform to weakly pyriform, 2-2.8 x 0.8-1.2 μm. Photobiont chlorococcoid. Spot tests: thallus K+ violet (section!), C- or C+ violet, KC-, P-; disc and proper margin of apothecia K+ red, C+ purple in section. Chemistry: thallus with the Sedifolia-grey pigment; disc and proper margin of apothecia with chlorinated anthraquinones.
Note: on isolated boulders of base-rich or lime-containing siliceous rocks, more rarely on compact limestone and dolomite. This taxon was included into C. conglomerata by Nimis (1993: 162). After having collected it in several parts of Italy, I now agree with Spanish authors in treating it as a distinct taxon, the parasitic habit and several morphological differences supporting this decision (see also Roux & coll. 2014).
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
paras Aspicilia spp.

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: very rare
Dry mediterranean belt: extremely rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29929)
2001/11/19



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0194504], Greece, East Aegean islands, Chios, Hill kefala, W along road Spartouda-Kambia, S-side of middle ridge, on sandstone rock. 38°33.87' N, 25°58.5' E, 480 m. Leg. H. Sipman (61370a) & T. Raus, 28. September 2013, det. H. Sipman, 2014.