Gyalolechia marmorata (Bagl.) Nimis & Arup

in Arup & al., Borziana, 4: 45, 2023. Basionym: Callopisma marmoratum Bagl. - N. Giorn. Bot. Ital., 11: 84, 1879
Synonyms: Caloplaca marmorata (Bagl.) Jatta non auct.; Caloplaca subochracea auct.; Caloplaca subochracea f. acrustacea Clauzade & Cl. Roux; Caloplaca subochracea f. pallida Clauzade & Cl. Roux; Caloplaca subochracea var. luteococcinea Clauzade & Cl. Roux
Distribution: C - Tosc (Nascimbene & al. 2021), Laz (TSB 9972), Sar (Arup & al. 2023). S - Camp (Aprile & al. (2003b), Pugl (Nimis & Tretiach 1999, Arup & al. 2023), Cal (Puntillo 1996), Si (Nimis & al. 1994, Ottonello & Salone 1994, Ottonello & al. 1994, Grillo & al. 2002, Grillo & Caniglia 2004, Caniglia & Grillo 2005, 2006, Genco & al. 2007, Arup & al. 2023).
Description: Thallus crustose, very variable, from endosubstratic and poorly evident to episubstratic and finely rimose-areolate, white to pale yellow, often delimited by a thin, black prothalline line. Apothecia biatorine, at first immersed, then sessile, rounded, 0.2-1.3 mm across, with a rusty red to dark orange-red, at first flat, then convex, epruinose disc, and a somewhat paler, smooth, at first well-visible, finally often excluded proper margin. Epithecium brownish orange, K+ red; hymenium colourless, 40-60(-80) µm high; paraphyses simple or forked in upper part, c. 1 µm thick at base, the apical cells (2-)3-4.5 µm wide; hypothecium colourless. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, ellipsoid, (9-)10-13(-14) x (3-)5-6(-10) µm, the equatorial thickening (“septum”) up to 5 µm. Photobiont chlorococcoid. Spot tests: thallus K- or K+ faintly red; C-, KC-, P-; Apothecia K+ red. Chemistry: thallus without lichen substances or with low amounts of fragilin; apothecia with parietin and emodin (major), and citreorosein, emodinal, fallacinal and parietinic acid (minor).
Note: a Mediterranean species found on compact limestone, locally very abundant in coastal situations, extremely rare far from the coast. The colour of the thallus, based on which several infraspecific taxa were distinguished (see e.g. Roux & coll. 2014); at least one of them is a distinct species.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Taxon bound to maritime-coastal situations

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: rather common
Dry mediterranean belt: rather rare

pH of the substrata:

12345

Solar irradiation:

12345

Aridity:

12345

Eutrophication:

12345

Poleotolerance:

0123

Altitudinal distribution:

123456


Predictive model
Herbarium samples

Michel Bertrand
France


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016


Courtesy: Olivier et Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_C/Textes_C5/Caloplaca_subochracea.htm
France, Sémaphore Pertusato, alt.100m - Corse
23/10/2016



P.L. Nimis CC BY-SA 04
TSB 36720



P.L. Nimis
holotype of Callopisma marmoratum Bagl. 1879



P.L. Nimis CC BY-SA 4.0
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis CC BY-SA 04
Herbarium: TSB 36720



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sardinia, Cagliari
09/06/2016
Holotype of Callopisma marmoratum Bagl (MOD)