Distribution: N - Frl (TSB 16892), TAA, Lomb, Piem (Morisi & Sereno 1995, Clerc & al. 1999, Isocrono & al. 2003), Emil (Fariselli & al. 2020), Lig. C - Marc (Nimis & Tretiach 1999). S - Camp (Garofalo & al. 2010). Bas (Herb. Ravera 1380), Cal (Puntillo 1996).
Description: Thallus squamulose, heteromerous, rather loosely attached, forming compact mats. Squamules 0.4-1.5 mm, usually convex, smooth, contiguous, ascending, imbricate, more or less lobulate, the peripheral ones often longer and somehow radiating; upper surface brownish grey to pale brown, greenish when wet, matt; lower surface black with a pale marginal zone, attached by dark rhizohyphae. Upper cortex pseudoparenchymatous; medulla white, rather lax. Perithecia half immersed in the squamules, 0.2-0.4 mm across, the upper part black, projecting around a depressed ostiole, without involucrellum. Exciple dark in upper part, pale in lower part; paraphyses absent, substituted by periphyses; hymenium I+ red, K/I+ blue; hymenial algae spherical to broadly ellipsoid. Asci (1-)2-spored, clavate to cylindrical-clavate, bi- and fissitunicate, thin-walled, the wall non-amyloid. Ascospores muriform, brown, elongate-ellipsoid, (25-)28-50 x (12-)14-22 µm. Pycnidia dark, immersed. Conidia crescent-shaped. 4-6 x 0.8 µm. Photobiont chlorococcoid (probably Diplosphaera, present in both thallus and hymenium). Spot tests: thallus and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly temperate, perhaps holarctic lichen found on terricolous mosses or on siliceous rocks, often near and on cyanobacterial colonies in periodically inundated sites, with optimum in upland areas with base-rich siliceous rocks.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris