Flavoplaca oasis (A. Massal.) Arup, Frödén & Søchting s.str.

in Arup & al., Nord. J. Bot., 31: 46, 2013. Basionym: Callopisma aurantiacum var. oasis A. Massal. - Sched. Crit., 7: 134, 1856.
Synonyms: Caloplaca aurantiaca f. oasis (A. Massal.) Th. Fr.; Caloplaca oasis (A. Massal.) Szatala
Distribution: N - VG, Ven (Lazzarin 2000b). C - Tosc, Marc (Nimis & Tretiach 1999), Umb (Genovesi & al. 2002, Ravera & al. 2006), Laz (Nimis & Tretiach 2004), Abr, Mol (Ravera & Genovesi 2012, Genovesi & Ravera 2014), Sar (Neuwirth 2018). S - Camp (Nimis & Tretiach 2004, Catalano & Aprile 2008, Garofalo & al. 2010), Pugl (Nimis & Tretiach 1999, Durini & Medagli 2004), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Grillo & al. 2007).
Description: Thallus crustose, endosubstratic to thinly and partially episubstratic, forming up to 1 cm wide, characteristic round patches on the thalli of endolithic Bagliettoa-species, the margins often discoloured and whitish, the centers orange-yellow, several patches tending to merge to cover much larger surfaces; the characteristic, round, discoloured patches are visible on the thalli of the host also when the lichen has died. Apothecia rather frequent, zeorine, scattered to crowded in the center of the patches, immersed to adnate, 0.2-04(-0.7) mm across, with a flat to slightly convex, orange disc, and a proper margin of irregularly radiating, thick-walled hyphae. Epithecium orange, granular, K+ red; hymenium and hypothecium colourless; paraphyses usually simple, sometimes branched above, 1.5-2.5 μm thick, the upper cells 4-5 μm wide. Asci 8-spored, cylindrical, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, ellipsoid, (7-)10-13(-15) x 5-7(-8) μm, the septum 3-7 μm thick. Photobiont: chlorococcoid. Spot tests: apothecia and pigmented parts of thallus K+ red, C-, KC-, P-. Chemistry: apothecia and sometimes thallus with parietin (major), fallacinal, emodin, teloschistin and parietinic acid (minor), corresponding with chemosyndrome A of Søchting (1997).
Note: a mild-temperate lichen found on hard, compact limestone in sites with plenty of diffuse light, such as in open deciduous forests; locally common in beech forests of southern Italy. This species has been much misunderstood (see Arup 2009 and Roux & coll. 2014): here I restrict the records to those referring to specimens forming small, rounded, characteristic islands on the thalli of Bagliettoa-species (especially B. calciseda, Roux in litt.) in natural habitats, while the non-parasitic forms occurring on limestone and concrete are provisionally dealt with under the following entry. In the past, this lichens might have been confused with F. polycarpa, and the Italian samples need revision
Growth form: Crustose endolithic
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
paras Bagliettoa spp.

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: rare
Montane belt: rather common
Submediterranean belt: rather rare
Padanian area: absent
Humid submediterranean belt: rather rare
Humid mediterranean belt: rare
Dry mediterranean belt: very rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm; Owner: CC BY-SA 4.0
[TSB_11707], Italia, Sicilia, Prov. Palermo, Madonie Mnts., loc. Portella Mandarini above Grerace Siculo, c. 1400 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & J. Poelt, 06.1988



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (32049)
2001/12/05
on the host, Bagliettoa parmigerella



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_11707], Italia, Sicilia, Prov. Palermo, Madonie Mnts., loc. Portella Mandarini above Grerace Siculo, c. 1400 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & J. Poelt, 06.1988



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_11707], Italia, Sicilia, Prov. Palermo, Madonie Mnts., loc. Portella Mandarini above Grerace Siculo, c. 1400 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & J. Poelt, 06.1988


Pier Luigi Nimis CC BY-SA 4.0
Herbarium: TSB 16669



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_11707], Italia, Sicilia, Prov. Palermo, Madonie Mnts., loc. Portella Mandarini above Grerace Siculo, c. 1400 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & J. Poelt, 06.1988



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_11707], Italia, Sicilia, Prov. Palermo, Madonie Mnts., loc. Portella Mandarini above Grerace Siculo, c. 1400 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & J. Poelt, 06.1988



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992.



Felix Schumm - CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992.



Felix Schumm - CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992.



Felix Schumm - CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992.



Felix Schumm - CC BY-SA 4.0
[TSB_16669], Italia, Friuli Venezia Giulia, Prov. Trieste, Gropada, near Abisso Plutone, on limestone in a doline, parasitic on Bagliettoa sp. Leg. M. Tretiach, 25.12-1992.



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_32392], Italia, Lazio, Prov. Latina, Aurunci Mountains at Monte Sant’Angelo near Gaeta, road to the top, c. 1200 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M. Tretiach, 05.04.2000



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Felix Schumm - CC BY-SA 4.0
[TSB_21965], Italia, Basilicata, Prov. Potenza, Viggiano, Acqua dei Pastori below Santuario di Madonna di Viggiano, 1370 m, on limestone, parasitic on Bagliettoa sp. Leg. P.L. Nimis & M.Tretiach, 11.04.1995.



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41386



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41386



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41386