Melanelia agnata (Nyl.) A. Thell

Nova Hedwigia, 60: 416, 1995. Basionym: Platysma agnatum Nyl. - Flora, 60: 562, 1877.
Synonyms:
Distribution: N - TAA (M 0024165).
Description: Thallus foliose, heteromerous, dorsiventral, closely adnate, forming up to to 10 cm wide patches. Lobes 0.5-2.5 mm wide, slightly concave to flat, with thicker, slightly up-turned margins. Upper surface dark brown to black (greenish brown in shade-forms), more or less shiny, with whitish, rounded to elongate, laminal and/or marginal pseudocyphellae; lower surface whitish to pale brown, darker along margins, somewhat wrinkled, with simple rhizines. Upper cortex paraplectenchymatous, with a non-pored epicortex, the cell walls containing isolichenan; medulla white. Apothecia common, lecanorine, laminal to submarginal, the disc concolorous with thallus, to 5 mm across. Epithecium brownish; hymenium and hypothecium colourless. Asci 8-spored, clavate, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid, 6.5-8.5 x 5-6.5 µm. Pycnidia common, black, sessile with a widened base. Conidia dumbbell-shaped (thickened at both ends), 5-7.5 x 1-1.5 µm. Photobiont chlorococcoid. Spot tests: upper cortex and medulla K-, C-, KC-, P-, N-, UV-. Chemistry: without lichen substances.
Note: a rather neglected, arctic-alpine and circumpolar species described from Italy and also known from Switzerland and France, growing on acid siliceous rocks near and above treeline. The type, collected by Arnold, is from the surroundings of Brenner Pass in South Tyrol.
Growth form: Foliose, broad lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: extremely rare
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

12345

Solar irradiation:

12345

Aridity:

12345

Eutrophication:

12345

Poleotolerance:

0123

Altitudinal distribution:

123456


Predictive model

Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC


LITERATURE


Acharius,EUnknown yearBemerkungen und Nachtrage zum Methodus Lichenum. Weber et Mohr Archiv fur die systematische Naturgeschichte

Acharius,EUnknown yearUsneae generis nova species. Nov. Act. regiae Soc. Scient. Upsaliensis

Alvarez, I/ Guzman-Davalos, L/ Guzman, GUnknown yearLiquenes su distribucion e importancia en Jalisco. Tiempos de Ciencia, Revista de Difusion Cientifica de la Universidad de Guadalajara

Martinengo, M/ Nimis, P/ Quaranta, L/ Durante, A (?) Licheni e Inquinamento.

Nair, AGR/ Subramanian, SSUnknown yearChemical components of Usnea flexilis Strtn.. Jour. Sci. & Indus. Res.

Re,G FUnknown yearAd Floram Pedemontanum Appendix.

Seaward, MRDUnknown yearLincolnshire psocids. Trans. Lincolnshire Nat. Union

Sipman, JHMUnknown yearStudies on Colombian cryptogams, X. The genus Everniastrum Hale and related taxa (Lichenes). Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, Series C

Stephenson, NL/ Rundel, PWUnknown yearQuantitative variation and ecological role of vulpinic acid and atranorin in the thallus of Letharia vulpina. Biochemical and Systematic Ecology

Weber, WAUnknown yearLichenes Exsiccati. Distributed by the University of Colorado. Fasc. 2 (no. 41-80).

Ruellius,J S (1536) De stirpium natura et historia libri XXX.

Dorstenius,T (1540) Botanicon continens herbarum, quarum usus in Medicina est, descriptiones et icones.

Gesner,C (1541) Historia plantarum et vires, ex Dioscoride, Aegineta, Theophr. Plinio et recentioribus Graecis juxta elementorum ordinem.

Fuchsius,L (1542) De stirpium historia commentarii insignes. http://library.si.edu/digital-library/book/dehistoriastirp00fuch+

Gesner,C (1542) Catalogus plantarum quadrilinguis.

Lonicerus,Adam (1551) Naturalis historiae opus novum.

Lacuna,Andr (1552) Commentaria in Dioscoridem cum figuris.

Tragus,Hyeronymus (1552) De stirpi libri tres.

Dodonaeus,Remb (1553) Stirpium historia Belgicae scripta et a Clusio Gallicae reddita, cum iconibus Antwerpiae.

Dodonaeus,R (1559) Stirpium historia Belgicae scripta et a Clusio Gallicae reddita, cum iconibus Antwerpiae.

Anguillara,A (1561) Opera botanica.

Cordus,V (1563) Sylva observationum variarum etc. edid. Conr. Gessner.

Mathiolus,P A (1565) Commentarii in libros VI Dioscoridis de materia medica.

Fallopius,G (1566) Opuscula a Petro Agatho edita.

Clusius,C (1576) De plantis per Hispaniam observatis libri duo, figuris 230 additis. Antwerpiae 1576.

Lobelius,M de (1576) Plantarum sive stirpium historia, cui annexum est adversarium volumen. Additis figuris 1476.

Lobelius,M de (1581) Plantarum s. stirpium icones.

Lonicerus,A (1582) Herbarium (Eucharii Roesleri).

Caesalpinus,A (1583) De plantis libri XVI.

Clusius,C (1583) De plantis per Pannoniam et Austriam observatis libri quatuor, cum fig. 353.

Mathiolus,P A (1583) Commentarii in libros VI Dioscoridis de materia medica.

Dodonaeus,R (1583-1616) Mechlinensis Medici caesarei, Stirpium Historiae Pemptades sex sive Libri XXX.

Castore,Durante (1585) Herbario nuovo.

Dalechampius,J (1586) Historia generalis plantarum. https://books.google.com/books?id=_NGRilRpGGIC&pg=PP4#v=onepage&q&f=false

Molinaeus,J (1587) Historia naturalis plantarum.

Camerarius,J (1588) De plantis Epitome utilissima Petri Andreae Mathioli Senensis.

Porta,J B (1588) Phytognomica VIII libris contenta.

Tabernaemontanus,J T (1588) Historia germanica tribus partibus edita cum fig. 2087.

Thal,J (1588) Sylva hercynica, seu Catalogus plantarum sponte nascentium in montibus et locis vicinis Hercyniae, qua respicit Saxoniam.

Wigandi,J (1590) Vera historia de Succino Prussiaco, de Alce Borussica et de herbis in Borussia nascentibus etc.

Lobelius,M de (1591) Plantarum s. stirpium icones.

Bauhin,C (1596) Phytopinax s. enumeratio plantarum nostro seculo descriptarum.

Gerardus,J (1597) Historia plantarum anglica.

Ferrantes,I (1599) Historiae naturalis lib. 28. cum fig.

Schwenkfeld,C (1600) Stirpium et Fossilium Silesiae Catalogus.

Clusius,C (1601) Historia plantarum rariorum cum appendice et auctuario et figuris 1135.

Lobelius,M de (1605) Observationes et Adversaria.

Columna,F (1606) Ecphrasis minus cognitarum rariorumque nostro coelo orientium stirpium.

Pilleter,C (1610) plantarum exoticarum et patriarum Walachriae synonymia alphabetico ordine praeposita. Middelburgi 1610.

Columna,F (1616) Ecphrasis minus cognitarum rariorumque nostro coelo orientium stirpium. Pars alt.

Bauhin,J (1619) Historiae generalis plantarum novae et absolutae Prodromus.

Bauhin,C (1620) Prodromus theatri botanici.

Bauhin,C (1623) Pinax theatri botanici sive Index in Theophrasti, Dioscoridis, Plinii et Botanicorum, qui a seculo scripserunt opera.

Johnson,T (1632) Enumeration plantarum in Ericato Hampstediano locisque vicinis crescentium.

Parkinson,J (1640) Theatrum botanicum or a herball of a large extent.

Bocconius,P (1647) Icones et Descriptiones rariorum Plantarum Siciliae, Melithae, Galliae et Italiae.

Paul,S (1648) Flora Danica. Hafniae 1648.

Bauhin,J (1650) Plantarum historia universalis nova et absolutissima etc.

Beniz Mart.,B de (1651) Observationes in Ephemeridibus nat. curios. Decas I.

Loesel,J (1654) Tractatum de plantis in Borussia sponte nascentibus.

Wormius,O (1655) Museum.

Rajus,J (1660) Catalogus plantarum circa Cantabrigiam nascentium.

Grisley,G (1661) Viridarium Lusitanicum. Ulyssipone 1661.

Rajus,J (1663) Appendix (to Catalogus Plantarum 1660).

Ambrosinus,H (1666) Phytologiae pars prima, Tom. 1.

Chabraeus,D (1666) Omnium stirpium Sciographia et Icones etc.

Merret,C (1667) Pinax rerum naturalium britann.

Borrichius,O (1670) De musco cathartico Islandiae. Actis Hafniensibus

Rajus,J (1670) Catalogus Plantarum Angliae. I

Breyne,J (1672) Miscell. Nat. Curios. Tom. III.

Pancovius,T (1673) Herbarium portatile per Barth. Zorn.

Scheffer,J (1674) Lapponia illustrata. Francof. 1674.

Hoffmann,M (1677) Florae Altdorffinae Diliciae sylvestres, sive Catalogus Plantarum in agro Altdorffino sponte nascentium.

Morison,R (1678) Historia plantarum universalis Oxoniensis. I

Wagner,J J (1680) Historia naturalis Helvetiae curiosa.

Mentzelius,C (1682) Index nominum plantarum multilinguis et pugillus rariorum plantarum.

Malpighi,M (1686) De plantis quae in aliis vegetant. Opera Omnia, Londoni

Plot,R (1686) Natural History of Staffordshire.

Rajus,J (1686) Historia plantarum, Vol. I.

Malphigius,M (1687) In Anatome Plantarum. 2. de plantis, quae in aliis vegetant.

Tabernaemontanus,J (1687) New vollkommentlich Krauterbuch.

Kyllingius,P (1688) Viridarium Danicum. Hafniae 1688.

Rajus,J (1688) Historia plantarum. Vol. II.

Bromelius,O (1694) Chloris Gothica seu Catalogus Stirpium circa Gothoburgum nascentium.

Rajus,J (1694) Stirpium extra Britanniam nascentium Sylloge.

Tournefort,J P de (1694) Elemens de botanique ou methode pour connaitre les plantes.

Cupani,F (1696) Hortus Catolicus.

Sloane,H (1696) Catalogus Plantarum Jamaicae.

Boccone,P (1697) Museo di Fisica e di esperienze, Parte I et Museo di Piante rare della Sicilia, Malta, Corsica etc. Parte II.

Magnolus,P (1697) Hortus regius Monspeliensis.

Petiver (1697-1703) Petiverani Musei Centuriae X.

Tournefort,J P de (1698) Histoire des plantes, qui naissent aux environs de Paris, avec leur usage dans la medicine.

Morison,R (1699) Historia plantarum universalis Oxoniensis. I

Buddlejus,A (1700) Hortus siccus plantarum Angliae, quo continetur curiosa Muscorum collectio.

Tournefort,J P de (1700) Institutiones rei herbarii. Edit. altera. (3 vols.).

Volkamer,J G (1700) Flora Noribergensis.

Gottsched (1703) Flora Prussica, sive Plantarum in regno Prussiae sponte nascentium.

Rajus,J (1704) Historia plantarum. Vol. III.

Sloane,H (1707) Historia naturalis jamaicae.

Scheuchzer,J J (1708) Itinera alpina tria.