in Zwackh, Lich. Exs.: 626, 626 bis (correction label), 1892.
Synonyms: Cladonia subcariosa auct. non Nyl.
Distribution: N - Frl (Tretiach & Hafellner 2000), TAA, Lomb (Rossi 1991, Rivellini & Valcuvia 1996, Gheza 2018, Isocrono & al. 2021), VA (Piervittori & Isocrono 1999), Piem (Ravera & al. 2017, Gheza 2018), Lig (Gheza & al. 2020). C - Abr (Nimis & Tretiach 1999, Gheza & al. 2021), Sar (Nöske 2000, Nöske & al. 2000).
Description: Primary thallus squamulose, persistent, the squamules large and prominent, ascending, forming dense mats, 3-10(-15) mm long, 2-5(-7) mm wide, olive-green above, white, but often becoming bluish, reddish brown or rarely brownish grey beneath. Podetia rather rare, cupless and thicker at apex, simple or rarely sparsely branched, often solid inside, olive-green to brown, 0.5-2 cm tall, c. 1.5 mm thick, almost continuously verruculose-corticate, esorediate, rarely squamulose at base. Apothecia common, dark brown, convex, forming larger subcorymbose aggregates which are 5-8 mm wide. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed on basal squamules, with a colourless jelly. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: K+ yellow rapidly turning red, C-, KC-, P+ yellow to red (slow reaction). UV-. Chemistry: norstictic acid, with traces of connorstictic acid, exceptionally homoheveadric acid, stictic or fumarprotocetraric acid complex, very rarely with atranorin.
Note: a mainly temperate lichen found on sandy mineral soil in open grasslands and on soil pockets on large isolated boulders, probably somehow overlooked in Italy and more widespread in the Alps.
Growth form: Fruticose
Substrata: soil, terricolous mosses, and plant debris