Clavascidium lacinulatum (Ach.) M. Prieto

in Prieto & al., Am. J. Bot., 99: 28, 2012. Basionym: Endocarpon hepaticum var. lacinulatum Ach. - Lichenogr. Univ.: 299, 1810.
Synonyms: Catapyrenium lacinulatum (Ach.) Breuss; Dermatocarpon trapeziforme auct. non (J. König) Trevis.; Placidium lacinulatum (Ach.) Breuss
Distribution: N - Ven, TAA. C - Sar. S - Pugl, Cal (Puntillo 1996).
Description: Thallus squamulose, pale to dark brown, dull, non- or very slightly pruinose. Squamules 3-8 mm broad, 0.3-0.5 mm thick, rounded at first, then crenulate, flattened to weakly convex, smooth, with an undulate-crisped edge, scattered to contiguous, rarely overlapping. Lower surface pale brown, sometimes darkening, attached by several, simple to weakly dichotomously branched, robust, 100-300 μm thick, several mm long, pale to brown rhizines, and by colourless, 3-5 μm thick rhizohyphae. Upper and lower cortex paraplectenchymatous, the lower cortex poorly developed, the cells not arranged in vertical rows; medulla at least partially of isodiametric, thick-walled cells, well-delimited from lower cortex. Perithecia frequent, laminal, immersed in the squamules, globose to broadly pyriform, up to 0.65 mm wide, without involucrellum. Exciple colourless to pale brown throughout; hymenium K/I+ blue; paraphyses absent, substituted by periphyses. Asci 8-spored, clavate, thin-walled, I-, without a tholus. Ascospores 1-celled, hyaline, ellipsoid, thin-walled, biseriate from the beginning, 12-15(-17) x 5.5-7.5 μm. Pycnidia laminal, immersed, black, to 0.5 mm in diam. Conidia oblong-ellipsoid, 3-4(-5) x 1.3-2 μm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly Mediterranean-Atlantic to mild-temperate terricolous species found on loess and calciferous ground, most frequent in dry grasslands at relatively low elevations.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: rare
Dry mediterranean belt: rare

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model
Herbarium samples

P.L. Nimis CC BY-SA 04
GZU Herb.Glowacki

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Glowacki

Pier Luigi Nimis - CC BY-SA 4.0
TSB 41278

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Glowacki