Paulia glomerata Henssen & Tretiach

Nova Hedwigia, 60: 298, 1995.
Synonyms:
Distribution: N - Frl (Henssen & Tretiach 1995, Tretiach & Molaro 2007).
Description: Thallus fruticose, black, gelatinous when wet, rarely orbicular, rosette-shaped and attached by a central holdfast, usually consisting of numerous, flattened or globose glomerules attached on conspicuous, elongated, whitish, branched, up to 3 cm long rhizoidal strands, the flattened lobes to c. 4 mm long and 2.8 mm broad, the globose ones to 2 mm across. Hyphae forming a network around the cyanobiont cells, which are penetrated by haustoria. Soredia abundant, especially at the base of the glomerules, globose, naked, 18-35 to 140 μm across. Apothecia rare, to 1.3 mm across, at first immersed then adnate, with a dark brown, urceolate to flat disc, and a c. 0.3 mm thick thalline margin. Proper exciple not developed; hymenium colourless, 120-180 μm high, non-amyloid; paraphyses sparingly branched and anastomosing; hypothecium colourless. Asci (2-)4-8-spored subcylindrical, without apical amyloid structures. Ascospores 1-celled, hyaline, ellipsoid to subglobose, 10-13(-17) x 9.5-11(-15) μm, the wall c. 1-2 μm thick. Pycnidia rare. Conidia bacilliform, 2.5-3.5 x c. 1 μm. Photobiont cyanobacterial, chroococcoid, the cells surrounded by a colourless to brown (in the peripheral parts of thallus) gelatinous sheath. Spot tests: all negative. Chemistry: without lichen substances.
Note: on steeply inclined surfaces of calcareous rocks in rainy areas, mostly starting its life-cycle in fissures of the rocks; to be looked for further in the Alps. The only Italian stations are from the Julian Pre-Alps, one of the rainiest areas in Europe.
Growth form: Fruticose

Substrata: rocks
Photobiont: cyanobacteria, coccaceous (e.g. Gloeocapsa)
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

12345

Solar irradiation:

12345

Aridity:

12345

Eutrophication:

12345

Poleotolerance:

0123

Altitudinal distribution:

123456


Predictive model
Herbarium samples


Felix Schumm; Owner: CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (16851)
2001/12/13
isotypus



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (16851)
2001/12/13
attachment organs, isotypus



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (16855)
2001/12/10



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (16855)
2001/12/10



Felix Schumm - CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015.



Felix Schumm - CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015.



Felix Schumm - CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015.



Felix Schumm - CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015.



Felix Schumm - CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015.



Felix Schumm - CC BY-SA 4.0
[TSB_16667], Italy, Friuli Venezia Giulia, Carnic Pre-Alps, Poffabro (PN) below Mt. Coltaront, c. 700 m on vertical limestone rocks. Leg. G. Bolognini & M. Tretiach, 18.04.1993; conf. M. Schulz, 02.2015.