Phaeophyscia ciliata (Hoffm.) Moberg
Symb. Bot. Upsal., 22, 1: 30, 1977. Basionym: Lichen ciliatus Hoffm. - Enum. Lich.: 69, 1784.
Synonyms: Lichen ulothrix Ach. non Hoffm.; Parmelia obscura var. ciliata (Hoffm.) Schaer.; Physcia ciliata (Hoffm.) Du Rietz; Physcia concrustans Nyl.; Physcia norrlinii Vain.; Physcia obscura auct. non (Ehrh.) Hampe ex Fürnr.; Physcia ulothrix (Ach.) Nyl.
Distribution: N - VG, Frl (Tretiach & Molaro 2007, Brackel 2013), Ven (Caniglia & al. 1999, Nascimbene & Caniglia 2003c, Nascimbene 2008), TAA (Nascimbene & al. 2006e, 2007b, 2022, Nimis & al. 2015), Lomb (Arosio & al. 2003, Brackel 2010, 2013, Gheza & al. 2020, 2023), Piem (Piervittori 2003, Isocrono & al. 2004, 2005b, 2007), VA (Piervittori & Isocrono 1999, Isocrono & Ongaro 2021, Ongaro & al. 2022), Emil (Fariselli & al. 2020), Lig (Brunialti & Giordani 2003). C - Tosc (Tretiach & Nimis 1994, Loppi & Frati 2006), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Ravera & al. 2006), Laz (Ravera 2001, Nimis & Tretiach 2004, Ruisi & al. 2005, Ravera & Genovesi 2008, Zucconi & al. 2013, Brackel 2015), Abr (Recchia & al. 1993, Olivieri & Pacioni 1996, Olivieri & al. 1997, 1997b, Loppi & al. 1999, Nimis & Tretiach 1999, Caporale & al. 2016, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Caporale & Ravera 2020), Sar (Rizzi & al. 2011, Di Nuzzo & al. 2022). S - Camp (Aprile & al. 2003, 2003b, Nascimbene & al. 2010b, Brunialti & al. 2013, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999), Bas (Bartoli & Puntillo 1998, Nimis & Tretiach 1999), Cal (Puntillo 1996, Diederich & al. 2010), Si (Campisi & al. 2020).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, closely adpressed, forming orbicular, rarely irregular, up to 5 cm (usually less) wide rosettes. Lobes elongate and discrete to (rarely) partly imbricate, 0.4-1.5 mm broad, flat, grey to brown-black in sun-forms, epruinose, sometimes with a continuous grey epinecral layer in older parts. Lower surface black, sometimes paler at lobe ends, with simple, black rhizines projecting beyond lobe margins. Upper and lower cortex paraplectenchymatous; medulla white. Apothecia frequent, lecanorine to 3 mm across (usually less), sessile, with a black, usually epruinose disc, and a thalline margin which often has a corona of dark rhizines in lower part. Epithecium brown; hymenium and hypothecium colourless; paraphyses slender, often forked in upper part, the apical cells clavate, with a thin dark cap. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, ellipsoid, 16-24(-27) x (7-)9-12(-13) µm, Physcia-type. Pycnidia black, immersed. Conidia ellipsoid, 3-3.5 x 1 μm. Photobiont chlorococcoid. Spot tests: K-, C., KC-, P-, UV-. Chemistry: without lichen substances. Note: a temperate to southern boreal lichen, most frequent on Fraxinus and Juglans in montane valleys, much less common than the closely related P. orbicularis, being absent from heavily disturbed areas and from eu-Mediterranean vegetation, and with narrower ecological requirements.
Growth form: Foliose, narrow lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather rare
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Austria: Alps (1100 m above sea level). On Sorbus aucuparia
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33628)
2001/12/07
Triin Aimla; Owner: University of Tartu
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Antonio Valero; Owner: Antonio Valero
Spain
2010
Antonio Valero; Owner: Antonio Valero
Spain
2010
Antonio Valero; Owner: Antonio Valero
Spain
2010
Antonio Valero; Owner: Antonio Valero
Spain
2010
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33628)
2001/12/07
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Austria: Alps (1100 m above sea level). On Sorbus aucuparia
Growth form: Foliose, narrow lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather rare
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Austria: Alps (1100 m above sea level). On Sorbus aucuparia
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33628)
2001/12/07
Triin Aimla; Owner: University of Tartu
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
Antonio Valero; Owner: Antonio Valero
Spain
2010
Antonio Valero; Owner: Antonio Valero
Spain
2010
Antonio Valero; Owner: Antonio Valero
Spain
2010
Antonio Valero; Owner: Antonio Valero
Spain
2010
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33628)
2001/12/07
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Felix Schumm - CC BY-SA 4.0
[20172], Austria, Tirolia, Samnaungruppe, Tösens, inter pagum Serfaus et capellam Sanctus Georgius, alt. 1300 m. Ad corticem Fraxini excelsioris. Leg. et det. J. Poelt et A. V zda, 7.09.1991. V ZDA: LICHENES SELECTI EXSICCATI NR. 2497
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Phaeophyscia_ciliata.htm
France
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
