Phaeophyscia hirsuta (Mereschk.) Essl.

Mycotaxon, 7: 302, 1978. Basionym: Physcia hirsuta Mereschk. - Ann. Cons. J. Bot. Genève, 121: 181, 1919.
Synonyms: Physcia labrata Mereschk. non sensu Frey; Physcia labrata var. olivacea Mereschk.
Distribution: N - VG (Castello 1996, 2002, Carvalho 1997, Martellos & Castello 2004, Castello & Skert 2005), Frl (Nimis & Salvadori 1998, Tretiach & Molaro 2007, Nascimbene & Salvadori 2008, Nascimbene & al. 2009b), Ven (Nimis & al. 1996c, Caniglia & al. 1999, Lazzarin 2000, Nascimbene & Salvadori 2008, Nascimbene 2008, Nascimbene & al. 2008e, 2015, Nascimbene & Marini 2010), TAA (De Benetti & Caniglia 1993, Nascimbene 2006c, 2014, Nascimbene & al. 2007b, 2014, 2022, Zarabska & al. 2009, Nimis & al. 2015), Lomb (Zocchi & al. 1997, Arosio & al. 2003, Furlanetto 2010), Piem (Caniglia & al. 1992, Piervittori 2003, Isocrono & al. 2005b, Furlanetto 2010, Giordani & Malaspina 2016), Emil (Gasparo & Tretiach 1996, Fariselli & al. 2020), Lig (Castello & al. 1994, Brunialti & al. 1999, Giordani & al. 2001, 2002, Brunialti & Giordani 2003, Giordani 2006, Benesperi & al. 2007, Giordani & Incerti 2008). C - Tosc (Tretiach & Nimis 1994, Loppi & al. 1994, 1995, 1997, 1997e, 1998, 1998b, 2002, 2002b, 2002c, 2003, 2004, Putortì & al. 1998, Benesperi 2000a, 2011, Del Guasta 2001, Lorenzini & al. 2003, Frati & al. 2006b, 2007, 2008, Brunialti & Frati 2010, Loppi & Nascimbene 2010, Paoli & al. 2012, 2012b, 2013, Brackel 2015, Nascimbene & al. 2015, Frati & Brunialti 2023), Marc (Frati & Brunialti 2006), Umb (Ravera 2000, Ravera & al. 2006), Laz (Ravera & al. 1999, Massari & Ravera 2002, Ruisi & al. 2005, Munzi & al. 2007, Ravera 2008b, Ravera & Genovesi 2008, Zucconi & al. 2013), Abr (Olivieri & al. 1997, 1997b, Loppi & al. 1999, Caporale & al. 2016, Corona & al. 2016), Mol (Frati & al. 2004, Caporale & al. 2008, Ravera & al. 2010, Paoli & al. 2011, 2015, Caporale & Ravera 2020), Sar (Rizzi & al. 2011, Di Nuzzo & al. 2022). S - Camp (Aprile & al. 2003b, Nascimbene & al. 2010b, Brunialti & al. 2013, Ravera & Brunialti 2013), Bas (Nimis & Tretiach 1999, Paoli & al. 2006), Cal (Puntillo 1996, Puntillo & Puntillo 2004, Incerti & Nimis 2006), Si (Grillo 1998, Grillo & Caniglia 2004, Ravera & al. 2023b).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, forming irregular to orbicular, up to 3 cm wide, often confluent rosettes. Lobes discrete to partly imbricate, 0.5-1(-1.5) mm broad, usually flat, grey to brown, with sparse to numerous, multicellular, transparent cortical hairs near the lobe ends, epruinose, with terminal, labriform soralia. Lower surface black, with simple rhizines. Upper and lower cortex paraplectenchymatous; medulla white. Apothecia rare, lecanorine, to 2 mm across, the thalline margin with cortical hairs. Epithecium brown; hymenium and hypothecium colourless; paraphyses slender, often forked in upper part, the apical cells clavate, with a thin dark cap. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, ellipsoid, 18-25(-28) x 7-12(-14) µm. Pycnidia rare, immersed. Conidia ellipsoid. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly temperate lichen found on isolated trees, more rarely on rock; most common in Tyrrhenian Italy. Some older records could refer to P. cernohorskyi.
Growth form: Foliose, narrow lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: rather rare
Humid mediterranean belt: rare
Dry mediterranean belt: extremely rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 04
TSB 42510



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (24288)
2001/12/03



Felix Schumm - CC BY.SA 4.0
[10270], Mexico, Sonora: 5 km west of Moctezuma, 109°44'W, 29°46'N, desert shrub and creek, on Prosopis, ca. 790 m Leg. T.H. Nash III (#34,071), 10.02.1993, det. T.H. Nash, 1993



Felix Schumm - CC BY.SA 4.0
[10270], Mexico, Sonora: 5 km west of Moctezuma, 109°44'W, 29°46'N, desert shrub and creek, on Prosopis, ca. 790 m Leg. T.H. Nash III (#34,071), 10.02.1993, det. T.H. Nash, 1993



Felix Schumm - CC BY.SA 4.0
[10270], Mexico, Sonora: 5 km west of Moctezuma, 109°44'W, 29°46'N, desert shrub and creek, on Prosopis, ca. 790 m Leg. T.H. Nash III (#34,071), 10.02.1993, det. T.H. Nash, 1993



P.L. Nimis CC BY-SA 04
TSB 42510



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (13672)
2001/12/04


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Phaeophyscia_hirsuta.htm
France, 11/10/2015 - région de Bonifacio, alt. 613 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Phaeophyscia_hirsuta.htm
France, 11/10/2015 - région de Bonifacio, alt. 613 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Phaeophyscia_hirsuta.htm
France, 11/10/2015 - région de Bonifacio, alt. 613 m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Phaeophyscia_hirsuta.htm
France, 11/10/2015 - région de Bonifacio, alt. 613 m - Corse


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=925&lang=en
France, Drôme Provençale