Punctelia subrudecta (Nyl.) Krog
Nord. J. Bot., 2: 291, 1982. Basionym: Parmelia subrudecta Nyl. - Flora, 69: 320, 1886.
Synonyms: Imbricaria borreri auct. ital. p.p.; Parmelia borreri auct.ital. p.p.; Parmelia borreri var. subrudecta (Nyl.) Clauzade & Cl. Roux; Parmelia dubia (Wulfen) Schaer.; Parmelia dubia f. scrobiculata B. de Lesd.; Parmelia dubia var. caesiocinerea B. de Lesd.; Parmelia helenae B. de Lesd.; Parmelia maculato-sorediosa (Gyeln.) Gyeln.; Punctelia helenae (B. de Lesd.) De Priest & B.W. Hale
Distribution: N - VG (Castello 1996, 2002, Leandrin 2004, Martellos & Castello 2004, Castello & Skert 2005, Thell & al. 2005, Leandrin 2006), Frl (Badin & Nimis 1996, Leandrin 2004, Castello & Skert 2005, Tretiach & Molaro 2007, Bernini & al. 2010), Ven (Nimis & al. 1996c, Nascimbene & Caniglia 1997, Caniglia & al. 1999, Lazzarin 2000, Valcuvia & al. 2000c, Leandrin 2004, Nascimbene 2005c, 2008, 2008c, Nascimbene & al. 2007, Nascimbene & Marini 2010), TAA (Philippi 1983, Gottardini & al. 2004, Leandrin 2004, Nascimbene 2006c, 2008b, 2014, Nascimbene & al. 2007b, 2014, 2022, Nimis & al. 2015), Lomb (Philippi 1983 Rivellini 1994, Arosio & Rinaldi 1995, Zocchi & al. 1997, Roella 1999, Arosio & al. 2003, Anderi & al. 2005, Valcuvia & Truzzi 2007b, Abramini & al. 2008, Furlanetto 2010, Gheza & al. 2015, 2020, Gheza 2019), Piem (Caniglia & al. 1992, Arosio & al. 1998, Piervittori 1998, 2003, Clerc & al. 1999, Isocrono & al. 2003, 2005b, Castino 2004, Leandrin 2004, Isocrono & Piervittori 2008, Furlanetto 2010, Matteucci & al. 2010, Giordani & Malaspina 2016), VA (Piervittori & Isocrono 1999, Valcuvia 2000, Valcuvia & al. 2000b), Emil (Bassi 1995, Gasparo & Tretiach 1996, Nimis & al. 1996, Sallese 2003, Leandrin 2004, Marconi & al. 2006, Morselli & Regazzi 2006, Cioffi 2009, Malavasi 2014, Gerdol & al. 2014, Fariselli & al. 2020), Lig (Castello & al. 1994, Adler & Ahti 1996, Putortì & al. 1999b, Giordani & al. 2002, Brunialti & Giordani 2003, Leandrin 2004, Giordani 2006, Giordani & Incerti 2008, Watson 2014). C - Tosc (Adamo & al. 1993, Loppi & Corsini 1995, 2003, Loppi & Putortì 1995, 1995b, Loppi & al. 1995, 1996, 1996b, 1996c, 1997, 1997e, 1998, 1998b, 1999a, 2002, 2002b, 2002c, 2003, 2004, 2004c, 2006, Adamo 1997, Monaci & al. 1997, Loppi & Nascimbene 1998, 2010, Putortì & al. 1998, Benesperi 2000a, 2006, Loppi & Pirintsos 2000, Bettini 2001, Del Guasta 2001, Lorenzini & al. 2003, Landi & Loppi 2003, Loppi & Frati 2004, Baruffo & Tretiach 2005, 2007, Frati & al. 2006b, 2007, Tretiach & al. 2008, Lastrucci & al. 2009, Brunialti & Frati 2010, Loppi & Baragatti 2011, Brackel 2015, Frati & Brunialti 2023), Marc (Gasparo & al. 1989, Nimis & Tretiach 1999, Leandrin 2004, Frati & Brunialti 2006, Brackel 2015, Nascimbene & al. 2015), Umb (Ravera 1998, Leandrin 2004, Ravera & al. 2006, Panfili 2007, Brunialti & al. 2012b, Brackel 2015), Laz (Bartoli & al. 1997, Ravera & al. 1999, Massari & Ravera 2002, Leandrin 2004, Nimis & Tretiach 2004, Ruisi & al. 2005, Munzi & al. 2007, Ravera 2008b, Zucconi & al. 2013, Brackel 2015), Abr (Recchia & al. 1993, Olivieri & al. 1997, 1997b, Loppi & al. 1999, Caporale & al. 2016), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Paoli & al. 2015), Sar (Loi & al. 2000, Zedda 2002, Rizzi & al. 2011, Di Nuzzo & al. 2022). S - Camp (Aprile & al. 2003b, 2011, Nimis & Tretiach 2004, Catalano & al. 2010, 2016, Nascimbene & al. 2010b, Garofalo & al. 2010, Brunialti & al. 2013, Ravera & Brunialti 2013), Pugl (Nimis & Tretiach 1999, Durini & Medagli 2004, Gianfreda & Matino 2020), Bas (Bartoli & Puntillo 1998, Nimis & Tretiach 1999, Leandrin 2004, Potenza 2006, Potenza & al. 2010), Cal (Puntillo 1996, Lich. Graec. 95: Obermayer 1997, Leandrin 2004, Puntillo & Puntillo 2004, Incerti & Nimis 2006, Brackel & Puntillo 2016, 2023), Si (Nimis & al. 1994, Ottonello 1996, Schicchi & al. 1997, Leandrin 2004, Caniglia & Grillo 2006b, Campisi & al. 2020, Ravera & al. 2023b).
Description: Thallus foliose, heteromerous, dorsiventral, forming 5-10 cm wide rosettes, sorediate. Lobes tightly adpressed, 5-10 mm wide, rounded and ascending at apices, grey or grey-green, with white, punctiform pseudocyphellae giving rise to white, orbicular, laminal soralia. Lower surface whitish to dark brown towards margins, pale brown towards center, with simple rhizines. Upper cortex paraplectenchymatous, of 2-6 layers of densely packed cells, with a non-pored epicortex, the cell walls with isolichenan; medulla white; algal layer continuous; lower cortex paraplectenchymatous. Apothecia very rare, lecanorine, 3-5 mm across, with a brown disc and a raised, often sorediate thalline margin. Proper exciple thin, colourless; epithecium brownish; hymenium and hypothecium colourless. Asci 8-spored, clavate, Lecanora-type. Ascospores 1-celled, hyaline, broadly ellipsoid, 14-18 x (7-)10-13(-15) µm. Pycnidia black, laminal. Conidia hook-shaped, 4-5 x c. 1 µm. Photobiont chlorococcoid. Spot tests: upper cortex K+ yellow, C-, KC-, P-; medulla and soralia K-, C+ red, KC+ red, P-, UV-. Chemistry: upper cortex with atranorin, medulla and soralia with lecanoric acid.
Note: a mainly temperate species (described, however, from the subantarctic Kerguelen Islands: the name in Europe may be misapplied!) found on bark of isolated deciduous trees, ecologically intermediate between Xanthorion and Parmelion; quite rare along the eastern side of the Peninsula and in the eu-Mediterranean belt, otherwise common below the montane belt.
Growth form: Foliose, broad lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: very common
Padanian area: rather rare
Humid submediterranean belt: very common
Humid mediterranean belt: rather rare
Dry mediterranean belt: very rare
Predictive model
Herbarium samples
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Sainte Anne la Palud
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31553)
2001/11/26
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31553)
2001/11/26
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli-Venezia Giulia, TS, Basovizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli-Venezia Giulia, TS, Basovizza; altitude: 2
2008.02.21
Harry Taylor; Owner: Natural History Museum London
Harry Taylor; Owner: Natural History Museum London
Harry Taylor; Owner: Natural History Museum London
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Conconello
14/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Punctelia_subrudecta.htm
France, 18/10/2018 - Ajaccio - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Punctelia_subrudecta.htm
France, 18/10/2018 - Ajaccio - Corse
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Peñas de Aia
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Growth form: Foliose, broad lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: very common
Padanian area: rather rare
Humid submediterranean belt: very common
Humid mediterranean belt: rather rare
Dry mediterranean belt: very rare
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Sainte Anne la Palud
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31553)
2001/11/26
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31553)
2001/11/26
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli-Venezia Giulia, TS, Basovizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli-Venezia Giulia, TS, Basovizza; altitude: 2
2008.02.21
Harry Taylor; Owner: Natural History Museum London
Harry Taylor; Owner: Natural History Museum London
Harry Taylor; Owner: Natural History Museum London
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Conconello
14/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Punctelia_subrudecta.htm
France, 18/10/2018 - Ajaccio - Corse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Punctelia_subrudecta.htm
France, 18/10/2018 - Ajaccio - Corse
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Le Faou
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Euskadi, Peñas de Aia
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=425&lang=en
Spain, Peñas de Aia
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Cap Sizun
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1387&lang=en
France, Drôme Provençale
fertile
