Pyrenodesmia alociza (A. Massal.) Arnold

Flora, 67: 310, 1884. Basionym: Biatorina alociza A. Massal. - Symmicta Lich.: 42, 1855.
Synonyms: Caloplaca alociza (A. Massal.) Mig.; Lecaniella alociza (A. Massal.) Jatta; Lecanora variabilis f. ecrustacea Nyl.; Placodium variabile var. ecrustaceum (Nyl.) Nyl.; Sporoblastia alocyza (A. Massal.) Trevis.
Distribution: N - VG (Crisafulli & al. 2006, Piervittori & al. 2006, Tretiach & Muggia 2006), Frl, Ven (Lazzarin 2000b, Tretiach & al. 2003, Nascimbene & Caniglia 2003c, Tretiach & Muggia 2006, Muggia & al. 2007, Thor & Nascimbene 2007), TAA (Spitale & Nascimbene 2012), Emil (Fariselli & al. 2020). C - Tosc (TSB 37172), Marc (Nimis & Tretiach 1999, Frolov & al. 2020), Umb (Nimis & Tretiach 1999, Ravera & al. 2006, Panfili 2007), Laz (Nimis & Tretiach 2004), Abr (Nimis & Tretiach 1999, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Genovesi & Ravera 2014), Sar. S - Camp (Aprile & al. 2003b, Nimis & Tretiach 2004, Garofalo & al. 2010), Pugl (Nimis & Tretiach 1999, Tretiach & Muggia 2006), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Nimis & al. 1994, Grillo 1998, Grillo & Caniglia 2004, Caniglia & Grillo 2005, 2006).
Description: Thallus crustose, endosubstratic or (rarely) very thinly episubstratic, white to grey, sometimes with a violet tinge, often delimited by a dark prothallus. Apothecia 0.2-0.8(-1) mm across, often slightly immersed in pits in the rock, with a flat to slightly convex, epruinose or slightly pruinose, black disc and a thin, finally excluded proper margin. Proper exciple brown in outer part, colourless to brownish within, the pigmented parts K+ violet; epithecium grey to brownish (in thick sections), K+ violet; hymenium colourless, with small crystals and with numerous extracellular oil droplets; apical cells of paraphyses 2-5 μm wide; hypothecium colourless. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, ellipsoid, (11-)15-18(-21) x (6-)7-8(-11) μm, the equatorial thickening (“septum”) 2.5-3(-5) μm. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-; pigmented parts of apothecia K+ violet and sometimes N+ pale violet. Chemistry: apothecia and sometimes thallus with the Sedifolia-grey pigment.
Note: on hard limestones and dolomite on sunny, exposed cliffs, with a wide altitudinal range; in Italy this is one of the most frequent species of Pyrenodesmia. For further details see Muggia & al. (2007).
Growth form: Crustose endolithic
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: extremely common
Montane belt: extremely common
Submediterranean belt: common
Padanian area: rare
Humid submediterranean belt: common
Humid mediterranean belt: rather rare
Dry mediterranean belt: rare

pH of the substrata:

12345

Solar irradiation:

12345

Aridity:

12345

Eutrophication:

12345

Poleotolerance:

0123

Altitudinal distribution:

123456


Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Domenico Puntillo; Owner: Domenico Puntillo
Italy



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15613)
2001/12/18



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



Felix Schumm - CC BY-SA 4.0
[2193], Jugoslavien, Sibenik, auf sonnigem Kalkgeröll bei Jadrija. Leg .et det. Schumm 1970, test. H. Wunder 1973.



P.L. Nimis CC BY-SA 04
TSB 15613


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_alociza.htm
France, session AFL août 2005 en Lozère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_alociza.htm
France, session AFL août 2005 en Lozère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_alociza.htm
France, session AFL août 2005 en Lozère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_alociza.htm
France, session AFL août 2005 en Lozère