Ramalina calicaris (L.) Fr.

Sched. Crit.: 17, 1824. Basionym: Lichen calicaris L. - Sp. Pl.: 1146, 1753.
Synonyms: Ramalina calicaris var. canaliculata Fr.; Ramalina calicaris var. evernioides (Anzi ex Jatta) Motyka; Ramalina fraxinea var. evernioides Anzi ex Jatta; Ramalina polymorpha var. crispa A. Massal. ex Beltr.
Distribution: N - Ven, TAA (Nascimbene & al. 2007b), Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020), Lig. C - Tosc, Marc, Laz, Abr, Mol (Garofalo & al. 1999, Nimis & Tretiach 1999, Caporale & al. 2008, Brackel 2020), Sar (Zedda 1995, 2002, 2002b, Zedda & al. 2001, Cossu 2013). S - Camp (Garofalo & al. 1999, Aprile & al. 2003b), Pugl (Garofalo & al. 1999, Nimis & Tretiach 1999, Durini & Medagli 2004, Brackel 2011), Bas (Brackel 2011), Cal (Incerti & Nimis 2006), Si (Brackel 2008b).
Description: Thallus greenish grey, fruticose, first tufted then pendent, to 20 cm long, stiff and hard when dry, the laciniae strap-shaped, little branched, 2-3 mm wide, typically gutter-shaped, the surface smooth with punctiform pseudocyphellae. Medulla solid. Apothecia usually frequent, marginal to subterminal, lecanorine, with a pale pink-ochre disc and a thin thalline margin. Asci 8-spored, clavate to cylindrical-clavate, the apical dome K/I+ dark blue with a pale, conical-pointed apical cushion (axial mass) never penetrating through the entire d-layer, the wall K/I-, but the thin outer gel K/I+ blue, Bacidia-type. Ascospores 1-septate, hyaline, straight, 10-16 x 5-7 µm. Photobiont: chlorococcoid. Spot tests: medulla K-, C-, KC-, P-. UV-. Chemistry: usnic acid in the cortex, sekikaic acid complex rarely present in the medulla.
Note: a mainly temperate species found on deciduous, more rarely coniferous trees, especially on branches in humid beech forests. Several earlier Italian records require confirmation, that from Venezia Giulia (see Nimis 1993: 596) was due to a misidentification.
Growth form: Fruticose

Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (9514)
2002/12/02



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (26707)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33012)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33012)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33012)
2008.03.10



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018



Felix Schumm - CC BY-SA 4.0
[WB877], Germany, Bayern, Oberbayern, Kreis Garmisch-Partenkirchen, Ammergebirge, Elmanbachtal, an Bergahorn, 1200 m, TK 8431/4. Leg. W.v. Brackel (no 8177), 12.7.2018
us: usnic acid


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Ramalina_calicaris.htm
France, 25/10/2016 - Site archéologique de Cucuruzzu, alt. 730m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Ramalina_calicaris.htm
France, 25/10/2016 - Site archéologique de Cucuruzzu, alt. 730m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Ramalina_calicaris.htm
France, 25/10/2016 - Site archéologique de Cucuruzzu, alt. 730m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Ramalina_calicaris.htm
France, 25/10/2016 - Site archéologique de Cucuruzzu, alt. 730m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Ramalina_calicaris.htm
France, 25/10/2016 - Site archéologique de Cucuruzzu, alt. 730m - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Ramalina_calicaris.htm
France, 25/10/2016 - Site archéologique de Cucuruzzu, alt. 730m - Corse


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Le Loc'h


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Le Loc'h


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Le Loc'h


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Le Loc'h


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Crozon


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France, Crozon


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=286&lang=en
France