Staurothele fissa (Taylor) Zwackh

Flora, 14: 552, 1862. Basionym: Verrucaria fissa Taylor in Mackay - Fl. Hibern., 2: 95, 1836.
Synonyms: Polyblastia umbrina (Wahlenb.) Rostr.; Sphaeromphale fissa (Taylor) Körb.; Sphaeromphale silesiaca A. Massal.; Staurothele hazslinszkyi (Körb.) Blomb. & Forssell; Staurothele inconversa (Nyl.) Blomb. & Forssell; Staurothele lithina sensu Zahlbr.; Staurothele silesiaca (A. Massal.) Zschacke; Staurothele umbrina (Wahlenb.) Hellb.; Thelotrema fissum (Taylor) Hepp; Verrucaria umbrina Wahlenb.
Distribution: N - TAA (Nascimbene 2005, Thor & Nascimbene 2007, Nascimbene & al. 2022), Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999). C - Marc, Sar (Nascimbene & al. 2023). S - Cal (Puntillo 1996).
Description: Thallus crustose, episubstratic, continuous to usually cracked, pale to dark brown, somewhat shiny, sometimes delimited by a whitish prothallus. Perithecia black, 0.2-0.3 mm across, immersed in 0.3-0.6 mm wide, conical-hemispherical projections with a brown to black apex, half to almost completely covered by a thalline layer. Involucrellum adpressed to exciple or slightly divergent in lower part, apical or extending to the base of the perithecium; hymenial gel I+ red (I+ blue at very low concentrations of iodine), K/I+ blue; hymenial algae subglobose to oblong-ellipsoid, 1-1.5 times as long as wide, 2-4(-5) μm; hamathecium of periphyses and periphysoids, interascal filaments absent. Asci 2-spored, clavate, the wall thickened above, with an ocular chamber, after dehiscence with a delicate extruded endotunica, I-, K/I-. Ascospores muriform, pale to dark brown, oblong-ellipsoid to narrowly oblong, (30-)37-50(-70) x 13-25(-30) μm. Pycnidia black, frequent, Dermatocarpon-type. Conidia rod-shaped or slightly curved, 3-4 x c. 1.2 μm. Photobiont chlorococcoid, present in both thallus and hymenium. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a probably circumpolar lichen growing amphibious in montane to Alpine creeks, or on moist siliceous rocks; much rarer in the high Mediterranean mountains than in the Alps.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Periodically submerged (e.g. in creeks)

Commonnes-rarity: (info)

Alpine belt: extremely rare
Subalpine belt: rather rare
Oromediterranean belt: extremely rare
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

12345

Solar irradiation:

12345

Aridity:

12345

Eutrophication:

12345

Poleotolerance:

0123

Altitudinal distribution:

123456


Predictive model
Herbarium samples


Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php (Courtesy: Anita Stridvall)



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Curtis Randall Björk CC BY-SA 4.0
British Columbia, Clearwater Valley Date: 2011-09-03 On siliceous boulder along fast-flowing river



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Sardinia
2014



Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
August 2017
as v. tilesii


Zschacke, H. (1934) Epigloeaceae, Verrucariaceae und Dermatocarpaceae. In: Dr. L. Rabenhorst‘s Kryptogamen-Flora, Band 9, Abt. 1, Teil 1. Akademische Verlagsgesellschaft, Leipzig, 695 pp. - Public Domain


Zschacke, H. (1934) Epigloeaceae, Verrucariaceae und Dermatocarpaceae. In: Dr. L. Rabenhorst‘s Kryptogamen-Flora, Band 9, Abt. 1, Teil 1. Akademische Verlagsgesellschaft, Leipzig, 695 pp. - Public Domain



P.L. Nimis CC BY-SA 04
TSB 10511


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1286&lang=en
Belgium, Ardennes