Toniniopsis verrucarioides (Nyl.) Kistenich, Timdal, Bendiksby & S. Ekman

Taxon, 676: 898, 2018.. Basionym: Lecidea aromatica var. verrucarioides Nyl. - Bot. Not.: 157, 1853.
Synonyms: Bilimbia carbonacea (Anzi) Jatta; Lecidea subimbricata Nyl.; Lecidea verrucarioides Nyl.; Thalloidima boissieri Müll. Arg.; Toninia aromatica var. cervina (Lönnr.) Th. Fr.; Toninia boissieri (Müll. Arg.) Arnold; Toninia carbonacea Anzi; Toninia cervina Lönnr.; Toninia congesta Hepp ex Kremp.; Toninia conjungens Th. Fr.; Toninia kolax Poelt; Toninia subimbricata (Nyl.) H. Olivier; Toninia verrucarioides (Nyl.) Timdal
Distribution: N - VG, Ven (Ravera & al. 2022b), Lomb, Lig (TSB 33570). C - Umb (Genovesi & al. 2002, Ravera & al. 2006), Sar. S - Cal (Puntillo 1996, Brackel & Puntillo 2016).
Description: Thallus squamulose, forming small patches to 1.5 cm diam., dark brown to dark grey, rarely weakly white-pruinose. Squamules scattered to partly imbricate, round to elongated, often crenulate, slightly convex, up to 2 mm wide. Upper cortex 50-120 μm thick, with an up to 70 μm thick epinecral layer; medulla white, lacking crystals. Apothecia frequent, lecideine, sessile, up to 1 mm across, black, epruinose or faintly pruinose, usually persistently flat and marginate. Proper exciple dark reddish brown, K-, N-; epithecium dark brown, sometimes with a faint green tinge, K-, N-, lacking crystals; hymenium colourless, 60-80 μm high; paraphyses not conglutinated, simple or sparingly branched and anastomosing in upper part, thin-walled, the apical cell distinctly swollen and covered by a gelatinous pigment cap; hypothecium dark reddish brown. Asci 8-spored, clavate, surrounded by a gelatinous I+ blue coat, with a well-developed I+ blue tholus, a I+ darker blue tube and a well-developed ocular chamber, Bacidia-type. Ascospores 1-3-septate, hyaline, ellipsoid to bacilliform, 10-19 x 3.5-5 μm. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-, UV-. Chemistry: thallus without lichen substances.
Note: an arctic-alpine to cool-temperate lichen found in fissures and fine crevices of calcareous rocks in upland areas, often growing on species of Placynthium when young.
Growth form: Squamulose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
paras Placynthium

Commonnes-rarity: (info)

Alpine belt: extremely rare
Subalpine belt: very rare
Oromediterranean belt: very rare
Montane belt: extremely rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

12345

(info)

Solar irradiation:

12345

(info)

Aridity:

12345

(info)

Eutrophication:

12345

(info)

Poleotolerance:

0123

(info)

Altitudinal distribution:

123456

(info)


Predictive model
Herbarium samples
Index Fungorum GBIF


P.L. Nimis CC BY-SA 04
TSB 27409



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33570)
2003/03/06
on Placynthium nigrum


Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC


Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine



Juri Nascimbene CC BY-SA 4.0
Italy, Veneto, Belluno, Vette Feltrine


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1149&lang=en
France, La Palue