Umbilicaria hyperborea (Ach.) Hoffm.

Deutschl. Fl., 2: 110, 1796. Basionym: Lichen hyperboreus Ach. - K. Vetensk.-Akad. Nya Handl., 15: 89, 1794.
Synonyms: Gyrophora aenea var. hyperborea (Ach.) Schaer.; Gyrophora hyperborea (Ach.) Ach.; Gyrophora hyperborea var. primaria Th. Fr.; Gyrophora ustulata (Vain.) Dalla Torre & Sarnth.
Distribution: N - Ven, TAA (Nascimbene & al. 2022), Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999). S - Bas (Potenza & al. 2014).
Description: Thallus foliose-umbilicate, dorsiventral, monophyllous, attached by a central holdfast, rigid and fragile, irregular in outline, (2-)4–6(-10) cm diam., deeply dissected, the edges down-turned, lacerated and often with oval perforations. Upper surface uniformly dark brown, sometimes paler towards the centre, covered with narrow, irregular, vermiform ridges often extending to margins and not forming a network. Lower surface uniformly dark brown, darker and almost black around umbilicus, paler towards the margins, smooth or occasionally dimpled, erhizinate or with very few rhizinomorphs, without thalloconidia. Upper cortex dark brown; medulla white, very loose under the ridges; lower cortex scleroplectenchymatous. Apothecia common, gyrodisc, black, sessile, mainly located between ridges over the whole upper surface except around the umbilicus, 1-1.5(-2) mm across. Asci 8-spored, clavate, thick-walled, with an amyloid apical dome, Umbilicaria-type. Ascospores 1-celled hyaline, ellipsoid, (7-)13-16(-20) x 3-8 µm. Photobiont chlorococcoid. Spot tests: K-, C+ red, KC+ red, P-. Chemistry: medulla with gyrophoric and umbilicaric acids, plus small amounts of lecanoric acid.
Note: an arctic-alpine, probably circumpolar lichen found on siliceous boulders wetted by rain, usually near the ground, with optimum near treeline; probably restricted to the Alps in Italy.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: rather common
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Mark Rahill - CC BY-SA 4.0
USA



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm – CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.



Felix Schumm - CC BY-SA 4.0
[19288], Australia, Victoria, Mt. McKay, Alpine National Park, Bogong High Plains, 16 km south-east of Mt. Beauty, 36°52' S, 147°14' E, 1840 m, on granite in alpine grasslands Leg. J.A. Elix (40558) & H. Streimann 18.02.1994. LICH. AUSTRALASICI EXS. 300.


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Frey E. 1933. Cladoniaceae (unter Ausschluß ded Gattung Cladonia) und Umbilicariaceae. In: Rabenhrst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz. 15: 1-208.


Giovanna Potenza
Italy, Basilicata, M. Volturino



P.L. Nimis CC BY-SA 04
TSB 13657


Ulrich Kirschbaum
Sweden: Dalarna.