Toninia tristis subsp. thalloedaemiformis (Szatala) Timdal

Opera Botanica, 110: 115, 1991. Basionym: Lecidea thalloedaemiformis Szatala - Borbásia, 3: 125, 1941
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Distribution:
Description: Thallus squamulose, chestnut brown to dark brown, shiny not forming cushions. Squamules contiguous to scattered, up to 4(-5) mm in diam., bullate, with punctiform impressions developing into pores, often forming cushions on rocks; lower surface pale. Upper cortex 40-160 µm thick, including an up to 100 µm thick epinecral layer, without crystals; algal layer continuous; medulla white, without crystals. Apothecia lecideine, sessile, strongly constricted at base, up to 2 mm across, with a black, flat to convex, smooth disc, and a distinct, smooth proper margin concolorous with disc. Proper exciple medium to dark brown (sometimes with a green tinge) in the rim, usually paler brown within, K-, N- or N+ violet, lacking crystals; epithecium olivaceous green to bright green, K-, N+ violet; hymenium colourless, 60-70 µm high; paraphyses not coherent, anastomosing, distinctly thickened above; hypothecium brownish. Asci 8-spored, clavate, surrounded by a gelatinous I+ blue coat, with a well-developed I+ blue tholus with a I+ darker blue tube and a well-developed ocular chamber, Bacidia-type. Ascospores 1-celled, hyaline, ellipsoid, 8-12 x 4-5 µm. Pycnidia dark, immersed. Conidia filiform. Photobiont chlorococcoid. Spot tests: thallus K-, C-, KC-, P-, UV-. Chemistry: thallus with terpenoids.
Note: this subspecies, which grows on steeply inclined calciferous rocks, has a mainly eastern Mediterranean distribution, being known from the Balkan Peninsula, Crete and the Peloponnesus, but is also known from the Southern Alps of France; to be looked for in Italy.
Growth form: Squamulose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

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Altitudinal distribution:

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Predictive model

Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC