Anema tumidulum P.M. Jørg., M. Schultz & Guttová

Henssen ex P.M. Jørg., M. Schultz & Guttová, Herzogia, 26: 2, 2013.
Synonyms:
Distribution: N - Frl, Ven (Nascimbene & al. 2021), Lomb (Jørgensen & al. 2013), VA (Matteucci & al. 2013). C - Tosc (Tretiach & al. 2008), Abr (Caporale & al. 2016).
Description: Thallus squamulose, homoiomerous, slightly gelatinous when wet. Squamules brown-black to black, dispersed to usually crowded to form effuse black-brown crusts, attached by a central point, to 3(-5) mm diam, shield-like, with an often lobulate to incised margin, not pruinose, often shiny, covered in granular isidia-like outgrowths which aggregate forming a dense bushy structure around the apothecia. Apothecia first immersed and perithecioid, but soon expanded, lecanorine, to 0.8(-1) mm across, with a dark reddish brown disc and a thick thalline margin. Hymenial gelatine I+ blue. Asci 4(-8)-spored, subcylindrical, prototunicate, thin-walled, with an amyloid apical cap. Ascospores 1-celled, hyaline, broadly ellipsoid to subglobose, 8-13(-17) x (4-)6-8(-11) µm. Pycnidia laminal, immersed. Conidia narrowly ellipsoid to bacilliform, c. 3 x 1 µm. Photobiont cyanobacterial, chroococcoid, of a few cells surrounded by a gelatinous sheath which is yellowish brown to purple near the thallus surface. Spot tests: all negative. Chemistry: no lichen substances.
Note: this species seems to be fairly common in central Europe (Jørgensen & al. 2013). It grows on steeply inclined, sunny surfaces of calcareous or basic siliceous rocks with periodical water seepage after rain, with optimum in upland areas. See also note on Anema moedligense.
Growth form: Crustose
Substrata: rocks
Photobiont: cyanobacteria, coccaceous (e.g. Gloeocapsa)
Reproductive strategy: mainly sexual
On otherwise dry surfaces with short periods of water seepage after rain

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather rare
Submediterranean belt: rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent


pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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P.L. Nimis CC BY-SA 04
TSB 34942