Gyalolechia bracteata (Hoffm.) A. Massal.

Ric. Auton. Lich. Crost.: 17, 1852. Basionym: Psora bracteata Hoffm. - Deutschl. Fl.: 169, 1796.
Synonyms: Caloplaca bracteata (Hoffm.) Jatta; Caloplaca bracteata f. alpina (Th. Fr.) Zahlbr.; Caloplaca bracteata f. deformis Erichsen?; Fulgensia bracteata (Hoffm.) Räsänen; Fulgensia bracteata subsp. deformis (Erichsen) Poelt?; Fulgensia bracteata var. alpina (Th. Fr.) Räsänen; Lecanora bracteata (Hoffm.) Röhl.; Placodium bracteatum (Hoffm.) Nyl.; Placodium fulgens var. alpinum Th. Fr.
Distribution: N - Frl, Ven (Nascimbene & Caniglia 1997, Caniglia & al. 1999, Brackel 2013), TAA, Lomb, Piem (Isocrono & al. 2004, Hafellner & al. 2004), VA (Piervittori & Isocrono 1999, Piervittori & al. 2001). C - Tosc (Vezda Lich. Rar. Exs. 172), Marc (Nimis & Tretiach 1999), Abr (Gheza & al. 2021, Di Nuzzo & al. 2021, Vallese & al. 2022).
Description: Thallus crustose, orange-yellow to golden yellow, of more or less dispersed groups of areoles, sometimes with poorly developed marginal lobes, forming 1-2 cm wide patches. Areoles thick, rounded, flat to mostly convex, scattered to (mostly) contiguous on a rather thick whitish hypothallus, not or only slightly dissected at margin, more or less pruinose, the marginal ones sometimes slightly elongate, to 1-2 mm long. Apothecia frequent, zeorine, sessile, 1-2 mm across, with an orange-brown, weakly concave to flat, smooth to rugose disc, and a thick, orange-yellow margin, which at maturity separates into a darker and thinner proper margin and a paler thalline margin. Epithecium brownish orange, K+ red; hymenium colourless, 60-70 µm high; hypothecium colourless. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 1-celled, hyaline, broadly ellipsoid, not restricted in the centre, thin-walled, 9-13(-15) x 4-7 µm. Pycnidia orange-yellow, immersed. Conidia narrowly ellipsoid. Photobiont chlorococcoid. Spot tests: thallus and apothecia K+ dark red, C-, KC-, P-, UV+ pale orange. Chemistry: fragilin dominant, with smaller amounts of emodin and parietin; physcion present in the apothecia.
Note: on calciferous soil and terricolous mosses in open situations, sometimes in fissures of calcareous rocks, mostly near or above treeline; parasitic of Thalloidima-species when young. The record of Grillo (1998) from the coast of Sicily, probably refers to G. subbracteata.
Growth form: Crustose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
paras Thalloidima sedifolium when young

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: rather common
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples
Index Fungorum GBIF


P.L. Nimis CC BY-SA 4.0
GZU Dupl.Graec.Lich. (59)



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (9283)
2001/12/24



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35718)
2003/01/22
sterile thallus


Andres Saag; Owner: University of Tartu



Curtis Randall Björk, - CC BY-SA 4.0
Near Hope Bay, Nunavut, Northwest Territories, Canadaries, CanadaIsland, British Columbia, Canada
19.07.2014



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Dupl.Graec.Lich. (59)
2003/04/28



Pier Luigi Nimis - CC BY-SA 4.0; Owner: Department of Life Sciences, University of Trieste
TSB 38006


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Textes_G1/Gyalolechia_bracteata.htm
France, Lanslebourg-Mont-Cenis, alt. 2000 m - Savoie
3/9/2013
var. alpina


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Textes_G1/Gyalolechia_bracteata.htm
France, Lanslebourg-Mont-Cenis, alt. 2000 m - Savoie
3/9/2013
var. alpina


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Textes_G1/Gyalolechia_bracteata.htm
France, Lanslebourg-Mont-Cenis, alt. 2000 m - Savoie
3/9/2013
var. alpina


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Textes_G1/Gyalolechia_bracteata.htm
France, Lanslebourg-Mont-Cenis, alt. 2000 m - Savoie
3/9/2013
var. alpina


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe: Germany.