Melanohalea elegantula (Zahlbr.) O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch
Mycol. Res., 108: 882, 2004. Basionym: Parmelia olivacea subsp. aspidota var. elegantula Zahlbr. - Verh. Ver. Nat. Heilk. Pressburg, 8: 39, 1894.
Synonyms: Collema exasperatum Ach.; Melanelia elegantula (Zahlbr.) Essl.; Melanelia incolorata (Parrique) Essl.; Melanohalea infumata sensu auct. medioeur. non (Nyl.) O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch; Parmelia elegantula (Zahlbr.) Szatala; Parmelia exasperatula var. elegantula (Zahlbr.) Zahlbr.; Parmelia incolorata (Parrique) Lettau; Parmelia jacquesii Werner
Distribution: N - VG (Castello 1996, Castello & Skert 2005), Frl (Tretiach & Molaro 2007), Ven (Nascimbene & Caniglia 1997, Caniglia & al. 1999, Nascimbene 2005c, Nascimbene & al. 2009c, Brackel 2013), TAA (Nascimbene 2003, Nascimbene & al. 2007b), Lomb (Rivellini 1994, Arosio & Rinaldi 1995, Zocchi & al. 1997, Roella 1999, Arosio & al. 2003, Stofer 2006, Brackel 2010, Furlanetto 2010, Gheza 2019b, Gheza & al. 2023), Piem (Morisi & Sereno 1995, Arosio & al. 1998, Isocrono & Falletti 1999, Piervittori 2003, Isocrono & al. 2004, 2007, Isocrono & Piervittori 2008, Furlanetto 2010, Matteucci & al. 2010, Giordani & Malaspina 2016), VA (Piervittori & Maffei 1996, Piervittori & Isocrono 1999, Valcuvia & al. 2000b, Matteucci & al. 2008, 2008c), Emil (Bassi 1995, Tretiach & al. 2008, Benesperi 2009, Gerdol & al. 2014, Fariselli & al. 2020), Lig (Brunialti & Giordani 2003). C - Tosc (Loppi & al. 1994, 1997, 1997b, 1998b, 2002, 2003, 2004c, 2006, Loppi & Putortì 1995b, Monaci & al. 1997, Loppi & Nascimbene 1998, 2010, Tretiach & Ganis 1999, Benesperi & al. 2007, Brunialti & Frati 2010, Benesperi 2011, Paoli & al. 2012, Brackel 2015, Frati & Brunialti 2023), Marc (Gasparo & al. 1989, Nimis & Tretiach 1999, Brackel 2015), Umb (Ravera 1998, Ravera & al. 2006), Laz (Ravera 2001, 2002, 2008b, Massari & Ravera 2002, Ruisi & al. 2005, Munzi & al. 2007, Ravera & Genovesi 2008, Zucconi & al. 2013, Brackel 2015), Abr (Nimis & Tretiach 1999, Brackel 2015, Caporale & al. 2016, Corona & al. 2016, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Ravera, Caporale & al. 2008, Genovesi & Ravera 2014, Brackel 2020, Caporale & Ravera 2020), Sar (Zedda 1995, 2002, Loi & al. 2000, Rizzi & al. 2011, Brackel & Berger 2019, Di Nuzzo & al. 2022). S - Camp (Aprile & al. 2002, 2003, Nimis & Tretiach 2004, Brunialti & al. 2010, 2013, Nascimbene & al. 2010b, Garofalo & al. 2010, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999, Brackel 2011), Bas (Nimis & Tretiach 1999, Fascetti & al. 2005, Potenza 2006, Potenza & al. 2010, Brackel 2011, Cassola & al. 2025), Cal (Puntillo 1995, 1996, Incerti & Nimis 2006, Stofer 2006, Brackel & Puntillo 2016), Si (Grillo & al. 1996, Grillo 1998, Grillo & Caniglia 2004, 2006, Falco Scampatelli 2005).
Description: Thallus foliose, heteromerous, dorsiventral, forming 4-7 cm wide rosettes, isidiate. Lobes (2-)3-4(-7) mm wide, contiguous to imbricate, adpressed but sometimes slightly ascending at margins, brown to olive-brown, usually dull, but sometimes shiny at lobe tips, epruinose, with abundant, cylindrical, simple to coralloid, 0.05-0.18 mm thick, not hollow isidia arising from conical papillae, with apical pseudocyphellae. Lower surface pale brown, slightly darker at margin, with mostly simple rhizines. Upper cortex paraplectenchymatous, with a non-pored epicortex, the cell walls containing isolichenan; medulla white; lower cortex paraplectenchymatous. Apothecia very rare, lecanorine, with a brown disc and an isidiate thalline margin. Epithecium brown; hymenium and hypothecium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid, 8-12 x 4-7 µm. Pycnidia rare, black, laminal. Conidia slightly dumbbell-shaped, c. 7 x 1 µm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, N-, UV-. Chemistry: without lichen substances.Note: a mild-temperate lichen found on old trees (e.g. oaks, Castanea), more rarely on siliceous rocks (Roux & coll. 2014), with optimum in the montane belt; common in the Apennines, rare in Northern Italy, and absent from urban areas. The saxicolous forms, that are worthy of further study, are not rare in the Alps, and were often confused with M. infumata, which is a strictly arctic-boreal species (see Esslinger 1977, Nimis & al. 2018).
Growth form: Foliose, broad lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather common
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
Herbarium samples
P.L. Nimis CC BY-SA 04
TSB 10984
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31232)
2001/12/21
tip of lobes
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28
Antonio Valero; Owner: Antonio Valero
Spain
2010
Pier Luigi Nimis - CC BY-SA 4.0
Herbarium: TSB 17796
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118
Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118
Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118
Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335
Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335
Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335
P.L. Nimis CC BY-SA 4.0
TSB 83637
P.L. Nimis CC BY-SA 4.0
TSB 83637
P.L. Nimis CC BY-SA 4.0
TSB 10984
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanohalea_elagantula.htm
France, Sur Genista salzmannii - Ajaccio - Corse
18/10/2018
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanohalea_elagantula.htm
France, Sur Genista salzmannii - Ajaccio - Corse
18/10/2018
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanohalea_elagantula.htm
France, Sur Genista salzmannii - Ajaccio - Corse
18/10/2018
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Thalkirchen
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Thalkirchen
Growth form: Foliose, broad lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather common
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
INDEX FUNGORUM
GBIF
DOLICHENS
P.L. Nimis CC BY-SA 04
TSB 10984
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31232)
2001/12/21
tip of lobes
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36850)
2008.02.28
Antonio Valero; Owner: Antonio Valero
Spain
2010
Pier Luigi Nimis - CC BY-SA 4.0
Herbarium: TSB 17796
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20156], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen, Straßenbaum an der Bergstraße, 48,73198° N, 9,63449° E, 368 m. Leg. et det. F. Schumm, 18.11.2018.
Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118
Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118
Felix Schumm - CC BY-SA 4.0
[20357], Austria, Stiria, Grazer Bergland, Graz, iuxta viam prope Wenisbuch, 550 m, in trunco Pyri communis. Leg. P. Döbbeler, J. Poelt & H. Teppner, 10.07.1972. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1118
Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335
Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335
Felix Schumm - CC BY-SA 4.0
[20356], Jugoslavia, Macedonia, montes Gali ica, secundum via inter Trpejca et Otešovo, 1550 m, ad cortices Quercuum vetustarum. Leg. A. Vezda, 11.07.1975. A. VEZDA: LICHENES SELECTI EXSICCATI NR. 1335
P.L. Nimis CC BY-SA 4.0
TSB 83637
P.L. Nimis CC BY-SA 4.0
TSB 83637
P.L. Nimis CC BY-SA 4.0
TSB 10984
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanohalea_elagantula.htm
France, Sur Genista salzmannii - Ajaccio - Corse
18/10/2018
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanohalea_elagantula.htm
France, Sur Genista salzmannii - Ajaccio - Corse
18/10/2018
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_M/Melanohalea_elagantula.htm
France, Sur Genista salzmannii - Ajaccio - Corse
18/10/2018
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1022&lang=en
France, Drôme Provençale
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 – Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Thalkirchen
