Ochrolechia androgyna (Hoffm.) Arnold
Flora, 68: 236, 1885. Basionym: Lichen androgynus Hoffm. - Enum. Lich.: 56, Tab. 7, f. 3, 1784.
Synonyms: Ochrolechia albosorediosa Gyeln.; Ochrolechia androgyna f. tatrica (Gyeln.) Verseghy; Ochrolechia androgyna var. albosorediosa (Gyeln.) Erichsen; Ochrolechia androgyna var. pergranulosa Räsänen; Ochrolechia androgyna var. saxorum auct. non (Oeder) Verseghy; Ochrolechia bahusiensis f. roseosorediosa (Gyeln.) Erichsen; Ochrolechia pulvinata var. ecorticata Verseghy; Ochrolechia roseosorediosa Gyeln.; Ochrolechia tartarea f. verrucosa Räsänen; Ochrolechia tartarea var. androgyna (Hoffm.) Arnold; Ochrolechia tatrica Gyeln.; Pertusaria degelii Erichsen; Pertusaria tumidula var. perpityrea Erichsen
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven (Nascimbene & al. 2005b, Nascimbene 2008c, 2011), TAA (Nascimbene 2006b, 2006c, 2008b, Nascimbene & al. 2006, 2007b, 2009, 2010, 2022, Thor & Nascimbene 2007, Zarabska & al. 2009, Kukwa 2011, Nimis & al. 2015), Lomb, Piem (TSB 33016), VA (Piervittori & Isocrono 1999, Valcuvia & al. 2000b, Piervittori & al. 2004), Emil (Fariselli & al. 2020). C - Tosc (Loppi & al. 1994, 1997, Loppi 1996, Loppi & De Dominicis 1996, Benesperi & al. 2007), Marc (Nimis & Tretiach 1999), Umb (Ravera & al. 2006b), Laz (Ravera 2001, Ravera & Genovesi 2008), Abr (Nimis & Tretiach 1999), Mol (Caporale & al. 2008, Paoli & al. 2015), Sar (Nöske 2000, Zedda 2002, 2002b, Zedda & Sipman 2001, Zedda & al. 2001, Rizzi & al. 2011, Neuwirth 2018, Di Nuzzo & al. 2022). S - Camp (Brunialti & al. 2013, Ravera & Brunialti 2013), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Liistro & Cataldo 2011, Campisi & al. 2020).
Description: Thallus crustose, superficial, thin to usually thick, smooth to bullate-tuberculate, white-grey to dark grey, delimited by a shiny, thick, pinkish grey prothallus, the tubercles usually not separated from one another, large, convex or flattened, in some specimens constricted at base. Soralia usually well delimited, 1.5-3 mm diam., round to irregular, rarely confluent and forming a continuous sorediate crust, concolorous with thallus or beige grey to yellowish- or pinkish-grey, bearing coarse granular, sometimes almost isidioid soredia. Cortex often covered with a distinct C– (necrotic) layer, white-grey to brown, often with a yellow tinge. Apothecia rare, lecanorine, 2-4 mm across, with a flesh-coloured, orange-brown, pale or dark brown, epruinose, at first concave then flat disc surrounded by a salmon-pink discoid tissue, and by a usually sorediate thalline margin. Thalline exciple corticate, with a well-developed medulla; proper exciple prominent, visible as a ring around thalline margin; hymenium colourless, up to c. 300 µm high; paraphyses thin, slender, densely branched and anastomosing; hypothecium colourless to yellowish. Asci 4-6(-8)-spored with thick, amyloid walls, without recognizable apical structures, Pertusaria-like. Ascospores 1-celled, hyaline, thin walled, broadly ellipsoid to subglobose, (12-)25-50 x (12-)17-31 µm. Photobiont chlorococcoid. Spot tests: cortex and epithecium K-, C+ pink, KC+ orange-pink, P-, UV-. Chemistry: gyrophoric acid (major), lecanoric acid (minor or trace), plus 3 fatty acids ('Androgyna B unknowns' sensu Tønsberg 1992). Note: on bark and on steeply inclined faces of siliceous rocks in humid montane forests, sometimes also on soil and bryophytes. The Italian material should be checked: some samples might belong to O. bahusiensis H. Magn. or to O. mahluensis Räsänen (see Kukwa 2011).
Growth form: Crustose
Substrata: rocks, bark, soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: very rare
Subalpine belt: rare
Oromediterranean belt: absent
Montane belt: rather rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

Predictive model
Herbarium samples


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36768)
2008.02.28


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36781)
2008.02.28


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36781)
2008.02.28


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_O/Ochrolechia_androgyna.htm
France, Bellefontaine - Jura
22/8/2016

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Crozon
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Crozon
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Pen-Hir
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Castel Meur
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Cap de la Chèvre
saxicolous, maritime

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Cap de la Chèvre
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime


Felix Schumm - CC BY 4.0
[2071], Schweiz, Merliwald, nördlich des Giswiler Stocks (651/187)
ca. 1350 m, an Südseite einer Abies, schattig. Leg. F. Schumm,
08.09.1972, det. F. Schumm, 1972. - Soredien: K-, C+ rot, P-
Growth form: Crustose
Substrata: rocks, bark, soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: very rare
Subalpine belt: rare
Oromediterranean belt: absent
Montane belt: rather rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

Predictive model
Herbarium samples |


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36768)
2008.02.28


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36781)
2008.02.28


P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36781)
2008.02.28


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form


Felix Schumm – CC BY-SA 4.0
8891], USA, Maine, Washington Co., Beals Island south of Jonesport, 44°29.348’ N, 67°35.664’ W, 32 m, Picea-forest, on granitic rock. Leg. Schumm 21.08.2001, det. F. Schumm, conf. E. Brodo 2013.
Unusual form

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_O/Ochrolechia_androgyna.htm
France, Bellefontaine - Jura
22/8/2016

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Crozon
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Crozon
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Pen-Hir
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Castel Meur
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Cap de la Chèvre
saxicolous, maritime

Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Cap de la Chèvre
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

Jacques Haine- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=730&lang=en
France, Pointe de Dinan
saxicolous, maritime

