Peltigera didactyla (With.) J.R. Laundon

Lichenologist, 16: 217, 1984. Basionym: Lichen didactylus With. - Bot. Arrang. Veget. Gr. Brit., 1, 2: 718, 1776.
Synonyms: Lichen caninus var. pellucidus Weber; Lichen spurius Ach.; Peltidea canina var. spuria (Ach.) DC.; Peltigera canina var. crispata J. Kickx f.; Peltigera canina var. notata Th. Fr.; Peltigera canina var. pusilla Fr.; Peltigera canina var. spuria (Ach.) Schaer.; Peltigera erumpens (Taylor) Lange; Peltigera leptoderma auct.; Peltigera pellucida (Weber) Gyeln.; Peltigera pusilla (Fr.) Körb.; Peltigera rufescens var. spuria (Ach.) Gyeln.; Peltigera rufescens var. vulnerata Müll. Arg.; Peltigera sorediata (H. Olivier) Fink; Peltigera spuria (Ach.) DC.; Solorina sorediifera Nyl.
Distribution: N - Frl (Tretiach & Molaro 2007), Ven (Nimis 1994, Caniglia & al. 1999, Nascimbene & Marini 2007), TAA (De Benetti & Caniglia 1993, Vitikainen 1994, Roux & Triebel 1994, Thor & Nascimbene 2007, Nascimbene 2008b, Lang 2009, Brackel 2013, Bilovitz & al. 2014, Nascimbene & al. 2022), Lomb (Gheza & al. 2022), Piem (E.C.I., 2, 843: Vitikainen 1994 Morisi & Sereno 1995, Isocrono & al. 2003, 2004, Morisi 2005, Isocrono & Piervittori 2008), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Matteucci & al. 2008c), Emil (Fariselli & al. 2020), Lig (Brunialti & al. 1999). C - Tosc (Vitikainen 1994), Marc, Laz, Abr (Nimis & Tretiach 1999), Sar. S - Camp (Jatta 1909-1911), Cal (Puntillo 1996), Si (Cataldo & Ravera 2012, Brackel 2008b).
Description: Thallus foliose, broad-lobed, heteromerous, dorsiventral, greenish grey to brownish when dry, blackish green when wet, forming 1-3(-5) cm wide rosettes or consisting of a few lobes only. Lobes 5-10 mm broad, rounded, concave, ear-shaped and sorediate when young, later ascending, esorediate and bearing vertically arranged apothecia. Soralia maculiform, laminar, flat, tending to disappear in fertile specimens (leaving discoloured patches), at first orbicular and 1-3(-5) mm across, then becoming confluent, with grey to grey-brown soredia. Upper surface tomentose, at least at margin; lower surface whitish, with narrow (0.4-0.7 mm broad) whitish to brownish veins and dense, up to 4 mm long rhizines becoming tufted and darkening. Upper cortex pseudoparenchymatous; medulla white; lower cortex absent. Apothecia rare in sorediate specimens, common in esorediate ones, saddle-shaped, brown, located at the tip of short ascending lobes. Paraphyses simple, distinctly thickened above. Asci fissitunicate, the thickened apex with a K/I+ blue ring, Peltigera-type. Ascospores 3-5(-7)-septate, hyaline to pale brown at maturity, fusiform, (36-)45-65(-80) x 3-5 μm. Pycnidia rare, dark, immersed. Conidia fusiform. Photobiont cyanobacterial (Nostoc, the cells not in long chains). Spot tests: cortex and medulla K-, C-, KC-, P- (soralia very rarely faintly C+ and KC+ pink). Chemistry: without lichen substances, or occasionally with low amounts of gyrophoric acid and methyl gyrophorate in soralia.
Note: a cool-temperate to boreal-montane, ephemeral lichen of disturbed mineral soil, most common in the Alps near and above treeline, becoming much rarer in Southern Italy.
Growth form: Foliose, broad lobed
Substrata: soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Pioneer species

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: rare
Oromediterranean belt: extremely rare
Montane belt: extremely rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Nere Sorrentino
Spain



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29377)
2001/12/10



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (29377)
2001/12/10
detail of soralia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (15055)
2003/03/03
lower surface


Triin Aimla; Owner: University of Tartu


Antonio Valero; Owner: Antonio Valero
Spain; altitude: 1900 m.
Herbarium: Forum Natura Mediterraneo
2011



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.



Felix Schumm – CC BY-SA 4.0
[13634], Germany, Baden-Württemberg, Kreis Göppingen, Rechberghausen nahe der Gärtnerei Epple, am Wegrand zwischen Gartenanlagen, 48°44.262' N, 9°38.505' E, 340 m, TK 7223. Leg. et det. Schumm 17.02.2008.


Nere Sorrentino
Spain


Mark Rahill - CC BY-SA 4.0
USA


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Hôpital-Camfrout


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=849&lang=en
France, Beaumont, Ardennes


Nere Sorrentino
Spain


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Nere Sorrentino
Spain