Peltigera horizontalis (Huds.) Baumg.
Fl. Lips.: 562, 1790. Basionym: Lichen horizontalis Huds. - Fl. Angl.: 453, 1762.
Synonyms: Lichen horizontalis var. nebulosus Vill.; Peltidea horizontalis (Huds.) Ach.; Peltigera horizontalis f. lacinulata Gyeln.; Peltigera horizontalis f. scabrida Oxner; Peltigera horizontalis var. muscorum Schaer.; Peltigera horizontalis var. pallidovenosa Savicz & Rass.; Peltigera horizontalis var. ticinensis De Not.; Peltigera horizontalis var. zopfii (Gyeln.) Trass; Peltigera ticinensis (De Not.) Jatta?; Peltigera zopfii Gyeln.
Distribution: N - VG, Frl (Vitikainen 1994, Nascimbene & al. 1998, Tretiach & Molaro 2007), Ven (Massalongo Lich. Ital. 214: Vitikainen 1994, Caniglia & al. 1999, Nascimbene & Caniglia 2003c, Nascimbene 2005c, 2008c, Nascimbene & al. 2009c, Nascimbene & Marini 2010, Nascimbene & al. 2013b), TAA (Vitikainen 1994, Nascimbene 2001b, 2006c, 2008b, Nascimbene & al. 2022), Lomb (Anzi Lich. Ital. 93 and Lich. Sondr. 60: Vitikainen 1994, Lang 2009, Brackel 2010, Gheza 2019, 2019b, Gheza & al. 2022, 2023), Piem (Vitikainen 1994, Triebel & al. 1997, Isocrono & al. 2003, Morisi 2005), VA (Vitikainen 1994, Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Matteucci & al. 2008c, Isocrono & Piervittori 2008), Emil (Nimis & al. 1996, Tretiach & al. 2008, Benesperi 2009, Brackel 2015, Fariselli & al. 2020), Lig (Vitikainen 1994, Brunialti & al. 1999). C - Tosc (Tretiach & Nimis 1994, Loppi & Putortì 1995b, Loppi & De Dominicis 1996b, Loppi & al. 1997b, 2004c, Putortì & al. 1999, Benesperi & Tretiach 2004, Benesperi 2006, 2011, Benesperi & al. 2007, Benesperi & Lastrucci 2007, Lastrucci & al. 2009, Nardini & al. 2013, Pasquinelli & al. 2013, Pasquinelli 2014, Brackel 2015), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Ravera & al. 2006, Panfili 2007), Laz (Vitikainen 1994), Abr (Nimis & Tretiach 1999, Caporale & al. 2016, Corona & al. 2016, Gheza & al. 2021), Sar. S - Camp (Garofalo & al. 2010, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999), Bas (Potenza 2006, Brackel 2011, Potenza & Fascetti 2012), Cal, Si.
Description: Thallus foliose, broad-lobed, heteromerous and dorsiventral, forming 10-30(-50) cm wide rosettes, bluish- to brownish grey when dry, blackish green when wet, loosely attached. Lobes 1-3 cm wide, up to 5 cm long, flattened, contiguous, with ascending margins, the upper surface glabrous, shiny, smooth. Lower surface pale towards margins, darker in central parts, with a very distinct pattern of rounded to oval, white patches in the spaces between the network of flat, dark brown-black, broad, reticulated veins, with fasciculate-squarrose, 2-3 mm long rhizines arranged in concentric lines. Upper cortex pseudoparenchymatous; medulla white; lower cortex absent. Apothecia frequent, terminal on short ascending lobes, with a brown, horizontal, flat disc. Paraphyses simple, distinctly thickened above. Asci 8-spored, fissitunicate, Peltigera-type, the thickened apex with a K/I+ blue ring. Ascospores 3(-5)-septate, hyaline, acicular, thin-walled, (25-)33-41(-47) x 3-7 µm. Pycnidia dark, immersed. Conidia fusiform. Photobiont cyanobacterial (Nostoc, the cells not in long chains). Spot tests: cortex and medulla K-, C-, KC-, P-, or medulla sometimes faintly and fleeting C+ and KC+ pale pink. Chemistry: two chemotypes, with different combinations of tenuiorin, methyl gyrophorate, gyrophoric acid, hopanes, and unidentified terpenoids.Note: on mosses (also epiphytic and epilithic) and humus-rich soil in the openings of humid forests, with a wide altitudinal range but with optimum in the montane belt.
Growth form: Foliose, broad lobed
Substrata: soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather common
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: extremely rare
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10582)
2001/12/05
apothecium
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34154)
2003/03/03
lower surface
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34154)
2003/03/03
rhizines
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33019)
2003/03/03
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010
Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start, 4: Position von Norstictinsäure, 7: Position von Atranorin, t: Tenuiorin, z: Zeorin
Juri Nascimbene – CC BY-SA 4.0; Owner: Department of ùLife Sciences, University of Trieste
Vinca, Prov. Massa Carrara, Tuscany, Italy
Mark Rahill - CC BY-SA 4.0
USA
Mark Rahill - CC BY-SA 4.0
USA
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, 4/12/2014 - Lans en Vercors - Servagnet - Isère
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, 4/12/2014 - Lans en Vercors - Servagnet - Isère
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Cher
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Cher
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Cher
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Huelgoat
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Doubs
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Doubs
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany; Baden-Württemberg: Black Forest.
Growth form: Foliose, broad lobed
Substrata: soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather common
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: extremely rare
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10582)
2001/12/05
apothecium
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34154)
2003/03/03
lower surface
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34154)
2003/03/03
rhizines
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33019)
2003/03/03
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010
Simonetta Peruzzi; Owner: Simonetta Peruzzi
Italy, Toscana, Firenze, Chianti
2010
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
Felix Schumm – CC BY-SA 4.0
[12539], Germany, Baden-Württemberg, Kreis Göppingen, Albrand, Bad Überkingen, auf dem Pfad zum Kahlenstein, 48°35.892' N, 9°48.493' E, 590 m, schattige Böschung. Leg. et det. Schumm 02.07.2006.
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start, 4: Position von Norstictinsäure, 7: Position von Atranorin, t: Tenuiorin, z: Zeorin
Juri Nascimbene – CC BY-SA 4.0; Owner: Department of ùLife Sciences, University of Trieste
Vinca, Prov. Massa Carrara, Tuscany, Italy
Mark Rahill - CC BY-SA 4.0
USA
Mark Rahill - CC BY-SA 4.0
USA
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, session AFL Luberon 2009 - Vaucluse
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, 4/12/2014 - Lans en Vercors - Servagnet - Isère
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_horizontalis.htm
France, 4/12/2014 - Lans en Vercors - Servagnet - Isère
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Cher
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Cher
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Cher
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Huelgoat
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Doubs
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Doubs
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=541&lang=en
France, Drôme
