Peltigera hymenina (Ach.) Delise

in Duby, Bot. Gall, 2: 579, 1830. Basionym: Peltidea hymenina Ach. - Meth. Lich.: 284, 1803.
Synonyms: Lichen polydactylus With. nom. illegit.; Peltigera lactucifolia auct. non (With.) J.R. Laundon; Peltigera polydactylon f. hymenina (Ach.) Flot.; Peltigera polydactylon var. crassoides Gyeln.
Distribution: N - Frl, TAA, Piem (Vitikainen 1994), Lig. C - Tosc (Loppi & al. 1994, 1997b). S - Camp (Catalano & al. 2016), Bas (Potenza 2006, Potenza & Fascetti 2012), Si (Alstrup 2004).
Description: Thallus foliose, broad-lobed, heteromerous and dorsiventral, grey to brownish when dry, blackish when wet, often somehow maculate, matt to glossy, loosely attached, forming up to 20 cm wide rosettes. Lobes elongate, flattened, contiguous, with rounded ends, 1-2 cm broad, with up-turned, undulate margins and a smooth, glabrous, upper surface. Lower surface largely pale ochre-coloured (darker only in central parts), with flat, broad, concolorous veins and pale, simple or fasciculate, up to 5 mm long rhizines. Upper cortex pseudoparenchymatous; medulla white; lower cortex absent. Apothecia frequent, saddle-shaped, terminal on short ascending lobes, brown. Paraphyses simple, distinctly thickened above. Asci 8-spored, fissitunicate, the thickened apex with a K/I+ blue ring, Peltigera-type. Ascospores 3-7-septate, hyaline turning pale brown at maturity, acicular, thin-walled, (47-)50-80(-90) x 3-5 µm. Photobiont cyanobacterial (Nostoc, the cells not in long chains). Spot tests: cortex and medulla K-, C-, KC-, P-, or medulla C+, KC+ pink. Chemistry: three chemotypes, with different combinations of tenuiorin, methyl gyrophorate, gyrophoric acid, peltidactylin, dolichorrhizin, traces of zeorin and of unidentified triterpenoids.
Note: on mineral soil in open, but never fully sun-exposed habitats, often associated with mosses, with optimum in the montane belt, from the Alps to the mountains of Southern Italy.
Growth form: Foliose, broad lobed
Substrata: soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rare
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Hôpital-Camfrout, Sanquer



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



P.L. Nimis CC BY-SA 04
TSB 34362



P.L. Nimis CC BY-SA 04
TSB 3151


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_hymenina.htm
France, session AFL 2006 - Finistère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_hymenina.htm
France, session AFL 2006 - Finistère


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P/Peltigera_hymenina.htm
France, session AFL 2006 - Finistère


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Plogonnec


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Plogonnec


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Crozon


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Plogonnec


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Rosnoen


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Rosnoen


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=210&lang=en
France, Rosnoen



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.



Felix Schumm – CC BY-SA 4.0
16644], Germany Rheinland-Pfalz, Burgruine Diemerstein bei Frankenstein, 49.444392°N, 7.961653° E, 275 m. Leg. Schumm 02.09.2010 (Blam-Exkursion), det. Schumm.


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany; Baden-Württemberg: Black Forest.