Lepra albescens (Huds.) Hafellner var. albescens
in Hafellner & Türk, Stapfia, 104: 171, 2016. Basionym: Lichen albescens Huds. - Fl. Angl., ed. 1: 445, 1762.
Synonyms: Marfloraea albescens (Huds.) S. Y. Kondr., L. Lőkös & J.-S. Hur; Pertusaria albescens (Huds.) M. Choisy & Werner; Pertusaria albescens f. globulifera (Turner) Ozenda & Clauzade; Pertusaria communis var. discoidea (Pers.) Garov.; Pertusaria communis var. variolosa (Flot.) Schaer.; Pertusaria dacica Erichsen; Pertusaria discoidea (Pers.) Malme; Pertusaria globulifera (Turner) A. Massal.; Pertusaria jurana Erichsen; Pertusaria leprarioides Erichsen non auct.; Pertusaria orbiculata (Schreb.) Zahlbr.; Pertusaria scutellata Hue; Pertusaria sorediata auct. ital.; Pertusaria tuberculata (Erichsen) Erichsen; Variolaria discoidea Pers.
Distribution: N - VG (Castello 1996, 2002, Carvalho 1997, Martellos & Castello 2004, Castello & Skert 2005, Craighero 2010), Frl (Badin & Nimis 1996, Craighero 2010), Ven (Lazzarin 1997, Caniglia & al. 1999, Nascimbene & Caniglia 2003c, Nascimbene & al. 2005b, 2006c, Nascimbene 2008c, Craighero 2010), TAA (Nascimbene & Caniglia 2000b, 2002c, Nascimbene & al. 2006e, 2007b, 2014, 2022, Thor & Nascimbene 2007, Zarabska & al. 2009, Craighero 2010, Nascimbene 2014, Nascimbene & Marini 2015, Nimis & al. 2015, Trindade & al. 2021), Lomb (Zocchi & al. 1997, Arosio & al. 2003, Stofer 2006, Valcuvia & Truzzi 2007b, Craighero 2010, Furlanetto 2010, Gheza & al. 2023), Piem (Caniglia & al. 1992, Arosio & al. 1998, Isocrono & Falletti 1999, Piervittori 2003, Isocrono & al. 2004, 2005b, 2006, Isocrono & Piervittori 2008, Craighero 2010, Furlanetto 2010, Matteucci & al. 2010, Lich. Ital. Exs. 26: Isocrono & al. 2019), VA (Piervittori & Isocrono 1999, Valcuvia & al. 2000b, Matteucci & al. 2008, Isocrono & al. 2008), Emil (Gasparo & Tretiach 1996, Nimis & al. 1996, Dalle Vedove & al. 2002, Tretiach & al. 2008, Benesperi 2009, Craighero 2010, Brackel 2015, Fariselli & al. 2020), Lig (Brunialti & al. 1999, Putortì & al. 1999b, Valcuvia & al. 2000, Giordani & al. 2002, Brunialti & Giordani 2003, Giordani 2006, Giordani & Incerti 2008, Craighero 2010, Watson 2014). C - Tosc (Tretiach & Nimis 1994, Loppi & Putortì 1995, 1995b, Loppi & al. 1995, 1997, 1997b, 1998, 1998b, 1999a, 2002, 2002b, 2002c, 2004c, 2006, Loppi 1995c, 1996, 1996b, Loppi & De Dominicis 1996, 1996b, Monaci & al. 1997, Loppi & Nascimbene 1998, 2010, Putortì & al. 1998, 1999, Tretiach & Ganis 1999, Putortì & Loppi 1999b, Bacci & al. 2000, Loppi & Pirintsos 2000, Benesperi 2000a, 2006, 2011, Laganà & al. 2002, Lorenzini & al. 2003, Loppi & Frati 2004, Frati & al. 2006b, Paoli & Loppi 2001, Benesperi & al. 2007, Lastrucci & al. 2009, Brunialti & Frati 2010, Craighero 2010, Paoli & al. 2012, 2012b, 2013, 2015d, Nascimbene & al. 2015, Frati & Brunialti 2023), Marc (Nimis & Tretiach 1999, Frati & Brunialti 2006, Craighero 2010, Brackel 2015), Umb (Ravera 1998, Nimis & Tretiach 1999, Panfili 2000b, 2007, Ravera & al. 2006, Craighero 2010, Brackel 2015), Laz (Ravera 2002, Massari & Ravera 2002, Ruisi & al. 2005, Ravera & Genovesi 2008, Craighero 2010, Zucconi & al. 2013, Brackel 2015), Abr (Recchia & al. 1993, Olivieri & Pacioni 1996, Loppi & al. 1999, Nimis & Tretiach 1999, Stofer 2006, Craighero 2010, Brackel 2015, Caporale & al. 2016, Corona & al. 2016, Gheza & al. 2021), Mol (Garofalo & al. 1999, Nimis & Tretiach 1999, Caporale & al. 2008, Craighero 2010, Ravera & al. 2010, Genovesi & Ravera 2014, Paoli & al. 2015, Brackel 2020, Caporale & Ravera 2020), Sar (Vezda Lich. Rar. Exs. 56, Zedda 1995, 2002, 2002b, Loi & al. 2000, Nöske 2000, Craighero 2010, Rizzi & al. 2011, Cossu 2013, Neuwirth 2018, Brackel & Berger 2019, Di Nuzzo & al. 2022). S - Camp (Garofalo & al. 1999, 2010, Ricciardi & al. 2000, Aprile & al. 2002, 2003, 2003b, Nimis & Tretiach 2004, Brunialti & al. 2010, 2013, Craighero 2010, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999, Durini & Medagli 2004, Craighero 2010, Brackel 2011), Bas (Nimis & Tretiach 1999, Craighero 2010, Brackel 2011), Cal (Puntillo 1995, 1996, Puntillo & Puntillo 2004, Incerti & Nimis 2006, Stofer 2006, Craighero 2010, Puntillo 2011, Brackel & Puntillo 2016), Si (Nimis & al. 1994, 1996b, Grillo & Cristaudo 1995, Grillo & al. 1996, 2002, Schicchi & al. 1997, Grillo 1998, Grillo & Caniglia 2004, Falco Scampatelli 2005, Caniglia & Grillo 2006b, Brackel 2008b, Craighero 2010, Liistro & Cataldo 2011, Campisi & al. 2020).
Description: Thallus crustose, episubstratic, continuous to fissured, whitish, pale grey or greenish grey, usually conspicuously zoned at margins, delimited by a white prothallus, not tasting bitter. Central parts of thallus with numerous, circular, 0.5-5 mm wide, usually concave to flattened, white soralia, the margin often somewhat raised; soredia white, granular (the forms corresponding to Pertusaria jurana have hemispherical, basally constricted, 0.5-1 mm wide warts, the largest ones with an orbicular soralium surrounded by a thin, raised thalline rim). Apothecia extremely rare, immersed in soralia-like warts and not visible without sectioning the warts, the disc densely white-pruinose. Hymenium colourless, the hymenial gel not or only weakly amyloid; paraphyses lax, branched and richly anastomosing. Asci 1(-2)-spored, broadly cylindrical, the apex without a distinct ocular chamber, the outer sheath K/I+ blue, otherwise K/I–, with an inner extensible layer, Pertusaria-type. Ascospores 1-celled, hyaline, ellipsoid, 170-300 x 50-115 µm. Photobiont chlorococcoid. Spot tests: thallus and soralia K-, C-, KC-, P-, UV-. Chemistry: allopertusaric (major) and dihydropertusaric (minor) acids. Note: a widespread, mainly temperate lichen found on bark, with optimum on old oaks, rare in agricultural areas (e.g. the Po-Plain) and in the Mediterranean belt, most common in deciduous open woodlands of the submediterranean and montane belts. On the synonymisation of Pertusaria jurana see Oset (2021).
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: very rare
Oromediterranean belt: absent
Montane belt: common
Submediterranean belt: rare
Padanian area: extremely rare
Humid submediterranean belt: rather common
Humid mediterranean belt: rare
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse. (Collected by Graf zu Solms; 1869! Identificated by Eichler/Cezanne).
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm - CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017.
Felix Schumm – CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017
Felix Schumm – CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017
Felix Schumm – CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017
Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain
Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain
P.L. Nimis CC BY-SA 4.0
TSB 38809
P.L. Nimis CC BY-SA 4.0
TSB 38809
P.L. Nimis CC BY-SA 4.0
TSB 41257
Andres Saag University of Tartu
P.L. Nimis CC BY-SA 04
TSB 9815
P.L. Nimis CC BY-SA 04
TSB 9815
Manuel Gil
Spain, Canary Islands, Tenerife
Manuel Gil
Spain, Canary Islands, Tenerife
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
acetone extract, crystals
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_4/Lepra_albescens.htm
France, sur écorce de frêne - session AFL 2010 - Hérault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Pyrenees, Azet
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Pyrenees, Azet
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Bagnoles de l'Orne
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Plougastel Daoulas
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Han-sur-Lesse
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Han-sur-Lesse
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Felix Schumm - CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse. (Collected by Graf zu Solms; 1869! Identificated by Eichler/Cezanne).
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: very rare
Oromediterranean belt: absent
Montane belt: common
Submediterranean belt: rare
Padanian area: extremely rare
Humid submediterranean belt: rather common
Humid mediterranean belt: rare
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse. (Collected by Graf zu Solms; 1869! Identificated by Eichler/Cezanne).
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm - CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017.
Felix Schumm – CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017
Felix Schumm – CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017
Felix Schumm – CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017
Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain
Erichsen C.F.E. 1936. Pertusariaceae. - In: Dr. L. Rabenhorsts Kryptogamenflora von Deutschland, Österreich und der Schweiz. Bd. IX, Abt. 5, T. 1. Akad. Verlagsges., Leipzig, pp. 321-512, 513-728. - Public Domain
P.L. Nimis CC BY-SA 4.0
TSB 38809
P.L. Nimis CC BY-SA 4.0
TSB 38809
P.L. Nimis CC BY-SA 4.0
TSB 41257
Andres Saag University of Tartu
P.L. Nimis CC BY-SA 04
TSB 9815
P.L. Nimis CC BY-SA 04
TSB 9815
Manuel Gil
Spain, Canary Islands, Tenerife
Manuel Gil
Spain, Canary Islands, Tenerife
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
acetone extract, crystals
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_4/Lepra_albescens.htm
France, sur écorce de frêne - session AFL 2010 - Hérault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Pyrenees, Azet
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Pyrenees, Azet
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Bagnoles de l'Orne
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Plougastel Daoulas
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Han-sur-Lesse
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=531&lang=en
France, Han-sur-Lesse
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1423&lang=en
France, Coadry
saxicolous
Felix Schumm - CC BY-SA 4.0
[19667], Germany, Baden-Württemberg, Kreis Heidenheim, östlich von Söhnstetten an der Abzweigung zum Dudelhof. 48.66927° N, 10.00898° E, 581 m. An alter Weidebuche in Trockenwiese-Hang. TK 7326. Leg. et det. F. Schumm, 07.08.2017.
