Lepra mammosa (Nyl.) Hafellner
in Hafellner & Türk, Stapfia, 104: 171, 2016. Basionym: Pertusaria mammosa Harm. - Lich. France, 5: 1141, 1913.
Synonyms: Marfloraea mammosa (Harm.) S. Y. Kondr., L. Lőkös & J.-S. Hur; Pertusaria etrusca Erichsen; Pertusaria mammosa Harm.
Distribution: C - Tosc, Laz, Sar (Nöske 2000). S - Cal (Puntillo 1996).
Description: Thallus crustose, episubstratic, 1-3(-4) mm thick, grey to grey-brown, sometimes zoned, markedly papillate, the papillae isidia-like, 1-2 mm wide, bearing at the top an initially punctiform, then expanded and 0.7-1 mm wide, white soralium with granulose soredia. Apothecia extremely rare, immersed on the top of fruiting papillae, with an expanded, dark grey to blackish disc. Hymenium colourless, the hymenial gel not or only weakly amyloid; paraphyses lax, branched and richly anastomosing; hypothecium colourless. Asci 1-spored, broadly cylindrical, the apex without a distinct ocular chamber, the outer sheath K/I+ blue, otherwise K/I–, with an inner extensible layer, Pertusaria-type. Ascospores 1-celled, hyaline, narrowly ellipsoid, 180-200 x 60-70 µm. Photobiont chlorococcoid. Spot tests: medulla and soralia K+ faintly yellow-brown, C-, KC+ reddish brown, P+ bright red. Chemistry: fumarprotocetraric acid, plus variable amounts of succinprotocetraric acid.Note: on steeply inclined to vertical surfaces of siliceous rocks in humid areas, usually below the montane belt; mainly Tyrrhenian in Italy.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: rare
Dry mediterranean belt: absent

Predictive model
Herbarium samples

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Harrie Sipman – Source http://www.bgbm.fu-berlin.de/sipman/Zschackia/AegeanLichens/CaloplacaAC.htm - As Caloplaca oasis

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_4/Lepra_mammosa.htm
France, session AFL 2010 - Hérault

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Cap Corse

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Col de Bavella

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: rare
Dry mediterranean belt: absent

Predictive model
Herbarium samples |

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Harrie Sipman – Source http://www.bgbm.fu-berlin.de/sipman/Zschackia/AegeanLichens/CaloplacaAC.htm - As Caloplaca oasis

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_L/Texte_L_4/Lepra_mammosa.htm
France, session AFL 2010 - Hérault

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Campomoro

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Cap Corse

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, Col de Bavella

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=917&lang=en
France, Corse, L'Ospedale
