Phlyctis agelaea (Ach.) Flot.
Bot. Zeit., 8: 574, 1850. Basionym: Lichen agelaeus Ach. - Lichenogr. Suec. Prodr.: 30, 1799.
Synonyms: Phlyctis italica Garov.; Phlyctis spilomatica (Schaer.) A. Massal. ?
Distribution: N - VG (Carvalho 1997), Frl, Ven, Lomb (Valcuvia 2002, 2002b, Gheza & al. 2023), Piem, Emil (Benesperi 2009, Fariselli & al. 2020), Lig (Giordani & Incerti 2008, Giordani & al. 2025). C - Tosc (Tretiach & Nimis 1994, Loppi & al. 1997b, Putortì & al. 1998, Paoli & Loppi 2001, Benesperi & al. 2007, Benesperi 2011), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Ravera & al. 2006, Panfili 2007), Laz (Bartoli & al. 1997, Ravera 2002, Ravera & al. 2003, Massari & Ravera 2002, Nimis & Tretiach 2004, Ruisi & al. 2005, Stofer 2006, Munzi & al. 2007, Zucconi & al. 2013), Abr (Nimis & Tretiach 1999, Stofer 2006, Caporale & al. 2016, Corona & al. 2016), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Ravera & al. 2010, Paoli & al. 2015, Caporale & Ravera 2020), Sar (Zedda 1995, 2002, 2002b, Loi & al. 2000, Vezda Lich. Rar. Exs. 448: Vezda 2000, Zedda & al. 2001, Rizzi & al. 2011, Cossu 2013, Neuwirth 2018, Di Nuzzo & al. 2022). S - Camp (Garofalo & al. 1999, 2010, Ricciardi & al. 2000, Aprile & al. 2003, 2003b, Nimis & Tretiach 2004, Brunialti & al. 2010, 2013, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999, Potenza 2006), Cal (Puntillo 1995, 1996, Puntillo & Puntillo 2004, Incerti & Nimis 2006), Si (Nimis & al. 1994, Grillo & Cristaudo 1995, Merlo 2004, Stofer 2006, Liistro & Cataldo 2011).
Description: Thallus crustose, ecorticate, thinly episubstratic, continuous or more or less cracked or verrucose, white to grey-white, forming up to 5(-12) cm wide patches delimited by a white prothalline line. Apothecia deeply immersed, clustered by 2-4 at the center of flattened, irregular, powdery or granular, 0.2-0.5(-1) mm wide thalline warts which are usually mistaken for soralia; disc concave to flat, brown-grey to reddish brown, but covered by the granular warts; thalline margin white-granular. Proper exciple thin, grey, I+ pale blue, usually obscured by the thalline exciple; epithecium colourless to brownish, strongly granular; hymenium colourless, K/I+ blue; paraphyses simple or sparingly branched in upper part, not capitate, c. 1.5 μm wide at base, c. 2.5 μm at apex; hypothecium pale brown. Asci 2(-4)-spored, broadly clavate, thin-walled, not strongly thickened at apex, the outer wall K/I+ blue, the tholus I+ weakly blue. Ascospores strongly muriform, hyaline to finally yellow-brown, broadly ellipsoid, clearly apiculate, (35-)45-80(-85) x (12-)14-30(-35) µm, with a thin, I+ blue wall and a very thin perispore. Pycnidia immersed, black. Conidia simple, ellipsoid, straight to rarely curved, 6-8 x 3-4 µm. Photobiont chlorococcoid. Spot tests: K+ yellow then slowly blood-red (needle-like crystals), C-, KC+ reddish, P+ yellow then orange-red, UV-. Chemistry: norstictic acid.Note: a mild-temperate to Mediterranean lichen found on acid-barked trees (especially Quercus ilex) in slightly sheltered but not very shaded situations; most frequent in Tyrrhenian Italy, much rarer in Northern Italy and along the eastern side of the Peninsula.
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: rare
Padanian area: absent
Humid submediterranean belt: rather common
Humid mediterranean belt: common
Dry mediterranean belt: extremely rare
Predictive model
Herbarium samples
Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11012)
2001/11/28
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11012)
2001/11/28
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Manuel Gil
Spain, Canary Islands, Tenerife
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 2012, session Normandie, gorges du Meillon, Le Vaudobin
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Landevennec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, St. Julien
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon
Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento
Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento
Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento
Felix Schumm - CC BY-SA 4.0
[VZR448], Italia. Insula Sardinia, Cagliari, reservatum naturae dictum
Monte Arcosu, prope domum forestalem Casa Perdu Molis, 500 m.
Ad corticem Quercus Ilicis. Leg. & det. A. Vezda, 29.8.1989. Ex A.
Vezda: Lichenes Rariores Exsiccati Nr. 448.
Felix Schumm - CC BY-SA 4.0
[VZR448], Italia. Insula Sardinia, Cagliari, reservatum naturae dictum
Monte Arcosu, prope domum forestalem Casa Perdu Molis, 500 m.
Ad corticem Quercus Ilicis. Leg. & det. A. Vezda, 29.8.1989. Ex A.
Vezda: Lichenes Rariores Exsiccati Nr. 448.
Felix Schumm - CC BY 4.0
[VZ1112], Romania. Distr. Hunedoara, Montes Retezat, in valle
Lapu ul Mare, supra Gura Apei, 1050 m. Ad corticem Aceris pseudoplatani.
Leg. A. Vezda, 5.6.1972. EX A. VEZDA LICHENES SELECTI
EXSICCATI NR. 1112.
Felix Schumm - CC BY 4.0
[VZ1112], Romania. Distr. Hunedoara, Montes Retezat, in valle
Lapu ul Mare, supra Gura Apei, 1050 m. Ad corticem Aceris pseudoplatani.
Leg. A. Vezda, 5.6.1972. EX A. VEZDA LICHENES SELECTI
EXSICCATI NR. 1112.
Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: rare
Padanian area: absent
Humid submediterranean belt: rather common
Humid mediterranean belt: common
Dry mediterranean belt: extremely rare
Predictive model
| Herbarium samples |
INDEX FUNGORUM
GBIF
DOLICHENS
Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11012)
2001/11/28
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11012)
2001/11/28
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Felix Schumm – CC BY-SA 4.0
[17975], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Nassach und Oberwälden, Waldweg zwischen Stärrerberg und Gaibhalden, 48.75491° N, 9.54289° E, 377 m, an Esche. Leg. Schumm 03.04.2012, det Schumm 2012.
Manuel Gil
Spain, Canary Islands, Tenerife
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 2012, session Normandie, gorges du Meillon, Le Vaudobin
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Phlyctis_agelaea.htm
France, 16/9/2017 - Cortambert - Saône-et-Loire
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Landevennec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Lesteven
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, St. Julien
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Tromel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Roscanvel
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=452&lang=en
France, Crozon
Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento
Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento
Wolfgang von Brackel - CC BY-4.0
Sporen - Italy, Cilento
Felix Schumm - CC BY-SA 4.0
[VZR448], Italia. Insula Sardinia, Cagliari, reservatum naturae dictum Monte Arcosu, prope domum forestalem Casa Perdu Molis, 500 m. Ad corticem Quercus Ilicis. Leg. & det. A. Vezda, 29.8.1989. Ex A. Vezda: Lichenes Rariores Exsiccati Nr. 448.
Felix Schumm - CC BY-SA 4.0
[VZR448], Italia. Insula Sardinia, Cagliari, reservatum naturae dictum Monte Arcosu, prope domum forestalem Casa Perdu Molis, 500 m. Ad corticem Quercus Ilicis. Leg. & det. A. Vezda, 29.8.1989. Ex A. Vezda: Lichenes Rariores Exsiccati Nr. 448.
Felix Schumm - CC BY 4.0
[VZ1112], Romania. Distr. Hunedoara, Montes Retezat, in valle Lapu ul Mare, supra Gura Apei, 1050 m. Ad corticem Aceris pseudoplatani. Leg. A. Vezda, 5.6.1972. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1112.
Felix Schumm - CC BY 4.0
[VZ1112], Romania. Distr. Hunedoara, Montes Retezat, in valle Lapu ul Mare, supra Gura Apei, 1050 m. Ad corticem Aceris pseudoplatani. Leg. A. Vezda, 5.6.1972. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1112.
